sample research proposal for phd in molecular biology

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Ph.D. Thesis Proposal

Procedures and policy guidelines, a. procedures.

The Graduate Advisory Committee (GAC) evaluates all Ph.D. Thesis Proposals. The GAC consists of at least seven (7) faculty members of the Department, and of one graduate student representative. Current GAC faculty members can be found online*.

• Graduate students must present their Thesis Proposals in the 5th or the 6th semester of their graduate studies.
• The Graduate Advisory Committee convenes twice a year to evaluate proposals (Fall and Spring).
• Proposals are usually scheduled near the end of semesters (December, May).

Before the Examination

• Graduate students must have passed (overall satisfactory) their yearly presentation in front of a Research Advisory Committee (RAC), with the more recent RAC no more than 6 months.
• Permission to present a Ph.D. Thesis Proposal is required and should be clearly stated in the most recent RAC report.
• The title of the Ph.D. Thesis Proposal must be submitted at least five (5) weeks in advance of the examination to Pascale Joseph ( undergrad.biochem [at] mcgill.ca ).
• The complete written Ph.D. Thesis Proposal must be submitted at least three (3) weeks in advance to Pascale Joseph.
• The written proposal must be prepared and formatted as described in Appendix A  below.
• The student must prepare a twenty (20) minute oral presentation of their Ph.D. The presentation should not be longer than 20 minutes. Thesis Proposal (PowerPoint or the like) as instructed in Appendix B below.

Thesis Proposal Examination

• Examinations are done in person and should take no more than 1 hour. This is usually conducted in room 908 of the Biochemistry Department.
• Before the start of the examination, the GAC will meet without the student to review the student’s progress (RAC reports, course performances, Junior Seminar evaluation), flag any areas of concern, and discuss the written Ph.D. Thesis Proposal.
• The student will deliver a 20-minute oral presentation of their Ph.D. Thesis Proposal. The overall question period should last no longer than approximately 40 minutes.
• The meeting outcome (satisfactory, conditional, or unsatisfactory)
• Identify any areas of concern and corrective actions
• Determine any other advice to be provided to the students
• Based on the outcome, set the date of the next meeting if necessary
• The GAC Chair will summarize the examination outcome in a letter to be submitted to the graduate student and supervisor.
• Graduate students must register to the BIOC702 (Ph.D. Thesis Proposal) course.

B.  Policy on Ph.D. Thesis Proposal Procedure and Outcome

The Ph.D. Thesis Proposal examination consists of the following three (3) components:

• The written Thesis Proposal . The written proposal must be prepared and formatted as per Appendix A. Incomplete or poorly written proposals will impact the GAC’s decision and examination outcome. Failure to submit the written proposal in due time may also impact the student’s status in the program.
• The oral presentation . The oral presentation should summarize the content of the written document and be prepared as outlined in Appendix B. The student should be able to defend their Thesis Proposal based on the presentation. Presentations that are poorly prepared or that last longer than 20 minutes will impact the GAC’s decision and examination outcome.
• Answers during the question period . General competence will be assessed in the question period. Students must be able to discuss and explain presented experiments and provide details as needed. They must be able to elaborate beyond what is shown in the presentation (e.g. provide alternative questions, experiments, analyses, interpretations). They must demonstrate some general knowledge outside of their immediate research by answering connected and general questions in their research fields. Failure to demonstrate competence could impact the GAC’s decision and examination outcome.

Each component of the exam will be rated as Satisfactory , Conditional , or Unsatisfactory . The overall outcome of the Ph.D. Thesis Proposal is determined as follows:

Satisfactory

When all 3 components are rated as Satisfactory, the student will be deemed to have completed all three components and will receive a “PASS” grade on the BIOC702 course (Ph.D. thesis Proposal). The next meeting will be with a RAC at their yearly presentation.

Conditional

If one or more of the components is rated as less than Satisfactory and no more than one component is Unsatisfactory , the student will be graded as “CONDITIONAL”. In the letter submitted to the student and supervisor following the examination, the GAC will impose one or more condition and a timeline for the student to fulfill them (normally no more than 3 months). If the student fulfills the conditions set by the GAC, the student will be awarded a “ Satisfactory ”, be given a “PASS” for BIOC702, and be granted permission to continue in the program. If not, the meeting outcome will be deemed “ Unsatisfactory ”, and the procedures outlined in the following section will be followed.

Unsatisfactory

When two (2) or more components of the exam are rated as Unsatisfactory, the overall meeting outcome will be graded as “UNSATISFACTORY”. In keeping with GPS regulations**, the following consequences will result:

In the case where a student is unsuccessful at the first attempt of the exam:

• A grade of “HH – to be continued” will be recorded for BIOC702
• The student will be required to repeat the entire exam at a time determined by the GAC. As per GPS regulations, this meeting must occur not sooner than 4 months and not later than 6 months after the first meeting.
• A student who is successful in the second attempt will be deemed to have passed the exam, and will be given a “PASS” for BIOC702.

In the case where a student is unsuccessful at the second attempt of the exam:

• The student will be deemed to have failed the Ph.D. Thesis Proposal.
• A grade of “FAIL” will be entered for BIOC702 replacing the previously recorded HH. The student will be required to withdraw from the program.

Preparing and formatting the written Ph.D. Thesis Proposal

The written Ph.D. Thesis Proposal should be prepared with the oral presentation in mind. This document should properly introduce the project, justify its importance, describe your research progress to date, and propose future experiments that will overall represent an important contribution to research. The Ph.D. Thesis Proposal should feature substantial preliminary experimental work and propose future experiments . The proposed experiments should be well thought of and feasible.

• Approximately and not more than 5 pages single spaced including figures.
• Figures should be inserted in line with the text, be legible, and not exceed 1 page of total space.
• References are not included in the document’s 5 page maximum.
• The main text should provide the following:
• Central hypothesis
• Specific Aims: these could represent your proposed thesis chapters. Three (3) proposed results Chapters is common but not required. In each of your Aims, specify the short and long term objectives.
• Significance
• You can include extra pages reporting additional work that does not fit in the main Thesis Proposal. This is optional and should be clearly marked as an Appendix.
• Document must be letter size (21.25 x 27.5 cm / 8.5 x 11 inches).
• In the header on the left, indicate your name (the lab you are in).
• In the header on the right, indicate the section title (Ph.D. Thesis Proposal)
• In the footer, include the page number (center) and date (right)
• A minimum margin of 2 cm all around the page is mandatory.
• Use a font size of 12 point. Single line spacing. No condensed type or spacing.
• Each section of your document should be properly separated (e.g. skip a line between sections, use bold font for section titles, etc.)
• Figures must have legends.

As per GPS ruling:

“The work submitted for this assessment is expected to be your own. The use of technologies such as ChatGPT are prohibited and will be considered a violation of the Code of Student Conduct.”

Preparing the Ph.D. Thesis Proposal presentation

The Ph.D. Thesis Proposal presentation should summarize the written proposal. It should contain everything needed to explain and defend the project and proposed experiments. With the presentation, you should be able to convince the Examiners that the Proposal is Satisfactory . Importantly , the Ph.D. Thesis Proposal is not merely a progress report: it should also clearly propose future experiments.

Your oral presentation should last approximately 20 minutes and not more. You can choose to have questions during your presentation or only after. The question period should last no more than 40 minutes.

The presentation should:

• Properly introduce the research question and project .
• Clearly state the main hypothesis, Aims and proposed experiments.
• Include substantial preliminary experimental work. You must clearly specify who did the presented experiments. Acknowledge the contribution of others where relevant. This includes figures or data published elsewhere, which must be accompanied by a citation.
• Clearly outline future directions and propose new experiments.

Evaluation criteria

• Quality and clarity of the presentation and it’s delivery.
• Quality of responses to questions about the design, background, and potential of the project. This includes experiments already performed and those suggested as future directions.
• Quality of responses to general background questions about biochemistry and molecular biology.
• Response to criticism raised during the Research Seminar 1 (Junior) examination.
• Feasibility of the proposed experiments.

Important note

• It is the responsibility of the student to inform themselves of the Ph.D. Thesis Proposal Procedures, Policy and deadlines. Failure to comply can lead to expulsion from the program, unless strong justification is provided and confirmed in writing by the Supervisor.
• The methods adopted for examination and evaluation are specified by Departmental regulations and are in agreement with the Graduate Studies guidelines approved by the Dean of Graduate and Postdoctoral Studies, McGill University.

Quick Guidelines

Before the examination:.

• Title of the proposal submitted five weeks in advance to Christine Laberge
• Complete written proposal submitted three weeks in advance to Christine Laberge
• Up to five pages single spaced
• Describe research progress to date, and proposed (future) research for the Ph.D. thesis
• Optional extra page indicating additional work not in the main proposal may be added and must be clearly marked
• Present the background, rationale, central hypothesis, specific aims and significance
• Include short and long term objectives
• Three proposed results chapters is common but not required
• Must include substantial preliminary experimental work

Thesis Proposal Examination:

• 20 minute presentation to GAC, 40 minute discussion
• Presentation summarizes the written proposal
• Questions about the design, background and potential of the project
• Questions about general background knowledge of biochemistry and molecular biology

Evaluation Criteria:

• Presentation, background knowledge and insight into the proposed research
• Knowledge in related areas of biochemistry
• Response to criticisms raised during the Research Seminar 1
• Feasibility of the proposed experiments

Revised: April 2023

*    Graduate Advisory Committee Members (GAC)

**  Graduate Student Supervision

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About the PhD in Biochemistry and Molecular Biology Program

In the Biochemistry and Molecular Biology PhD program, faculty, and students work together to increase knowledge of the biochemical and molecular bases of normal and abnormal cellular processes. Our program trains students to be successful independent scientists and gives them the knowledge, research training, and leadership skills to continue to provide new insights into the biomedical issues that have a profound impact on public health. Cancer biology is a historical and continuing area of interest for many faculty in our program, which has been supported in part by a training grant from the National Institutes of Health’s National Cancer Institute since 1975.

Students engage in a rigorous course curriculum and a range of structured and informal activities outside the classroom and lab to build their skills. They will pursue their thesis research in the lab of one of our over forty training faculty across the Johns Hopkins Bloomberg School of Public Health and the Johns Hopkins School of Medicine.

Visit our dedicated PhD program website to learn more about the diverse research training opportunities of the program.

PhD in Biochemistry and Molecular Biology Program Highlights

Our position within the School of Public Health provides a unique setting in which students learn how biochemistry, molecular biology, physical chemistry, cell biology, and genetics can be used to solve significant problems in public health and medicine. Our program offers:

• Training faculty from across the School of Public Health and the School of Medicine
• A strong grounding in the science of biomedical and public health research through a core curriculum that includes courses taught by leading experts from the Schools of Public Health and Medicine
• Training outside the lab and classroom in key skills such as communications and leadership
• Opportunities to build strong communications skills through a range of speaking venues including journal club, research colloquium, department retreats, and national meetings
• Teaching Assistant service to build teaching and interpersonal skills, with options for additional training and professional development through the Johns Hopkins University Teaching Academy to further develop skills
• Access to the Johns Hopkins School of Medicine Professional Development and Career Office , offering excellent career services and professional development,  including the BMB-required OPTIONS program, a guided process of career exploration for paths from medicine to biotech to academia and beyond
• Opportunities to participate in community service and outreach, with a focus on our East Baltimore neighborhoods, through the Johns Hopkins University community engagement and service-learning center, SOURCE

Training faculty across the School of Public Health and the School of Medicine

Schools that students can take courses in: Public Health, Arts & Sciences, Medicine, and Engineering

Two-month rotations in the first year prior to selecting thesis lab

Average number of incoming students in the BMB PhD degree program each year

What Can You Do With a PhD In Biochemistry And Molecular Biology?

The Biochemistry and Molecular Biology PhD program prepares students for a range of biomedical and health sciences careers, including in academia, industry, policy, and beyond. Visit the Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries.

Sample Careers

• Research Scientist
• Science Policy Adviser
• Biotech Executive
• Senior Scientist
• Patent Lawyer
• Science Policy Analyst/Advocate
• Science Writer/Journalist
• Biological Sciences Teacher

Topic Areas

The BMB PhD program faculty conduct research to gain new insights into the cellular and molecular mechanisms underlying normal and abnormal cellular processes, and their relevance as targets for improving health and treating disease. Our training program places particular emphasis on mechanistic approaches to research problems, and cancer biology has had a prominent place in our research interest for over 50 years .

Common topic areas within our faculty's diverse research interests include:

• Biophysics and Structural Biology
• Cancer Biology
• Chemical Biology and Proteomics
• Cell Biology
• Cellular Stress and Cell Signaling
• Genetics, Genomics, and Gene Regulation
• Immunology and Infectious Diseases
• Translational Research

Curriculum for the PhD in Biochemistry and Molecular Biology

The BMB PhD offers students a rigorous course curriculum, including a set of common core classes from the Schools of Public Health and Medicine. Students further tailor their curriculum with elective courses chosen based on interests and career goals, with options that span the Schools of Public Health, Medicine, Engineering, and Arts and Sciences. A rich array of seminar programs and journal clubs are also available to all students.

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Admissions Requirements

For the general admissions requirements see our How to Apply page. The specific program also requires:

Prior Research Experience

Laboratory research experience (from academia, industry, etc.) is required

Prior Coursework

Strong background in the sciences, particularly in chemistry, biochemistry, or biology

Standardized Test Scores

Standardized test scores (GRE) are optional for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Program Faculty Spotlight

Ashani T. Weeraratna

Ashi Weeraratna, PhD, studies how cancer cells move to distant sites and how changes in the normal cells around a tumor contribute to their movement, especially as we age.

Michael J. Matunis

Michael Matunis, PhD, studies how protein modification by SUMO—the small ubiquitin-related modifier—drives changes in key cellular pathways from stress response to DNA repair.

Jennifer M. Kavran

Jennifer Kavran, PhD, MS, MPhil, is a biophysicist who investigates how cells communicate with each other and their environment.

Danfeng Cai

Danfeng Cai, PhD, combines advanced microscopy, genomics, and proteomics to tease out the functions of protein condensates in cells, with a focus on cancer.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2024. VTSI applicants are eligible for an application fee waiver , but the fee waiver must be requested by November 15, 2024 and prior to submission of the SOPHAS application.

Per the Collective Bargaining Agreement (CBA) with the JHU PhD Union, the minimum guaranteed 2025-2026 academic year stipend is $50,000 for all PhD students with a 4% increase the following year. Tuition, fees, and medical benefits are provided, including health insurance premiums for PhD student’s children and spouses of international students, depending on visa type. The minimum stipend and tuition coverage is guaranteed for at least the first four years of a BSPH PhD program; specific amounts and the number of years supported, as well as work expectations related to that stipend will vary across departments and funding source. Please refer to the  CBA to review specific benefits, compensation, and other terms.

In the BMB PhD program, all full-time PhD students who remain in good academic standing will receive the above support through the entire duration of the program.

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU starting in Fall 2023 or beyond can apply to receive a need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help.

Mike Matunis, PhD PhD Program Director

Roza Selimyan , PhD BMB Executive Director for Academic Affairs and Education Programs

Erika Vaitekunas Administrative Specialist

[email protected]

We further basic science knowledge at biochemical and molecular levels that is relevant to public health.

News Programs Research Highlights Faculty

Biochemistry and Molecular Biology Headlines

BMB Professor Michael Matunis Receives 2024 Shikani/El-Hibri Prize for Discovery and Innovation

The award recognizes Matunis' research on SUMO and a recent discovery of its role in protein quality control. 

Exploring Cancer’s Ancestral Links

BMB Assistant Professor Brittany Jenkins-Lord seeks clues in the lab and the community to understand the biological causes of cancer disparities.

Weeraratna Lab Research Highlight

New paper shows age-related stiffening of skin can alter growth and barrier function of blood vessels in melanoma tumors, contributing to metastasis

What We Do in the Department of Biochemistry and Molecular Biology

Our Department strives to increase knowledge of cellular processes—normal and abnormal—at biochemical and molecular levels. We train highly qualified scientists who, through research, teaching, and service, continue to provide new insights into biomedical issues with a profound impact on public health.

Biochemistry and Molecular Biology Highlights

Founding department.

In 1917, we were the Department of Chemical Hygiene—our name has changed but not our drive for cutting-edge research.

Basic lab science with a public health perspective

Our research labs are uncovering the fundamental biochemical and molecular mechanisms at the foundations of health.

Teaching and training focus

We are committed to providing our students with training and mentoring so they can excel in a challenging but supportive environment.

From Cancer to CRISPR

Our faculty research—and teach—diverse topics across the breadth of biochemistry and molecular biology.

Biochemistry and Molecular Biology Programs

As a basic research department within the #1 public health school, the Department of Biochemistry and Molecular Biology thrives at a unique intersection of basic science, medicine, and public health.

We offer a doctoral and two master’s degree programs. We are also part of an online interdepartmental Master of Arts in Public Health Biology.

Master of Science (ScM)

A two-year program providing training in cellular and molecular biology, blending coursework in the first year with extensive lab research experience

MHS program focused on cellular and molecular biology at the intersection of biology, medicine, and public health, preparing students for a wide range of careers

Our students are immersed in cutting-edge research in biochemistry and molecular biology, providing insights into biomedical issues impacting public health

Ashi Weeraratna, PhD,

studies how cancer cells move to distant sites and how changes in the normal cells around a tumor contribute to their movement, especially as we age.

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Department news.

Support Our Department

A gift to our department can help provide student scholarships and internships, attract and retain faculty, and support innovation.

PhD projects 2024

New PhD projects for 2025 will be published soon. Stay tuned!

[2024] - Co-evolution of transposable element activity and host genome

[2024] - Understanding the Distribution of Mutations along Genomes

[2024] - Chromatin regulation in stem cells and development

[2024] - Molecular mechanisms of genome transcription regulation & dysregulation

[2024] - Investigating Antisense Oligonucleotide (ASO) Treatments for Neurodegenerative Diseases

[2024] - Transcriptional condensates

[2024] - Computational systems medicine and disease control

[2024] - Comparative Analysis of DNA Methylome Conservation Across Species

[2024] - Genome Regulation Department

[2024] - Synthetic biology of long-range gene regulation

[2024] - lncRNAs in 3D – dissecting the gene regulatory function of long non-coding RNAs 

[2024] - Evolution of primate transcription factor genes

[2024] - Mathematical modelling of cis-regulatory landscapes

[2024] - Epigenetic mechanisms controlling cell fate decisions during the early stage of liver, pancreas and biliary tree development

[2024] - Transcriptional Regulation Group

[2024] - Molecular mechanisms of bacterial immunity

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Ph.D. Program Biomolecular Structure and Mechanism

Quicklinks und sprachwechsel, main navigation, research proposal guidelines.

BSM Program, Life Science Zurich Graduate School Research proposal guidelines The format of the research proposal for the BSM PhD Program generally follows the proposal format required by the Department of Biology, ETH Zurich. Research proposals in the BSM PhD program should meet certain criteria regarding form and content. Excluding title page they should have a maximum length of five pages (about 2000 words) and be built up as follows: Title page • working title of the proposed dissertation • name and contact information of the doctoral student • Institute, at which the dissertation should be carried out • names of the members of the doctoral thesis committee • proposed dates for the beginning and planned end of the dissertation Content (approx. 2000 words, max. 5 pages) •  Introduction  (suitable for ‘general biologists’) – approximately  2 pages • descriptions of the  Questions/Hypotheses/Goals  that the dissertation should address:  ½ page •  Experimental approach/strategy: ½ page •  Results obtained  in the period until submission of the proposal:  max. 2 pages • a schematic  Timeline  (schedule of what should be achieved when and by which means) • a list of relevant  References  (cited in the text) At the end of the proposal (outside the 5-page limit), please also outline briefly: • teaching responsibilities • courses taken or intended to be taken for credit

Graduate School

Molecular Biology

General information, program offerings:, director of graduate studies:, graduate program administrator:.

The graduate program in the Department of Molecular Biology fosters the intellectual development of modern biologists. We welcome students from a variety of educational backgrounds and offer an educational program that goes well beyond traditional biology. The molecular biology department at Princeton is a tightly knit, cohesive group of scientists that includes undergraduate and graduate students, postdoctoral fellows, and faculty with diverse but overlapping interests. Graduate students have a wide choice of advisers, with a broad spectrum of interdisciplinary interests and research objectives.

The graduate program offers all entering students the opportunity, with the help of faculty advisers, to design the intellectual program that best meets their unique scientific interests. Each student chooses a series of research rotations with faculty members in molecular biology and associated departments (chemistry, computer science, ecology and evolutionary biology, chemical and biological engineering, physics, and neuroscience). Entering students, with the aid of the graduate committee, select core and elective courses from a large number of offerings in a variety of departments and disciplines. This combination of a cohesive department, one-on-one advising, and individualized programs of course work and research provides an ideal environment for graduate students to flourish as independent scientists.

Areas of concentration include biochemistry, biophysics, cancer, cell biology, computation and modeling, development, evolution, genetics, genomics, microbiology and virology, policy, and structural biology.

Program Offerings

Program offering: ph.d..

By the end of the first year, students must have completed four core courses, achieving an average of B or better. Students may take additional elective courses that are closely related to their research topic.

By the end of the third year, students must have completed MOL 561 - Scientific Integrity in the Practice of Molecular Biology.

Additional pre-generals requirements

Rotations Students must complete three laboratory rotations with different advisers as part of their research training during the first year of study (MOL 540, MOL 541 Research Projects); a fourth rotation is optional. Students who complete a full rotation (approximately 8 weeks of research) the summer before entering graduate school are assigned a rotation in September along with other entering students. A student may elect to work with any faculty member or associated faculty member of the program. Students who desire to work with faculty outside the program may do so only with the approval of the director of graduate studies.

Faculty/Student Research Talks In the fall of their first year, graduate students attend a series of informal talks given by each faculty mentor. These discussions are designed to introduce first-year students to current research projects that might serve as rotation and thesis topics.

Molecular Biology Annual Retreat The molecular biology annual retreat is a two-day symposium of research talks and poster sessions held in the fall and attended by all graduate students, postdoctoral fellows, and faculty in molecular biology and associated departments. 

Mol Bio Colloquia Graduate students, together with postdoctoral fellows and faculty, attend weekly research seminars given by graduate students and postdoctoral fellows. This colloquium provides both experience in the presentation of research results and a forum for scientific discussion with peers.

General exam

The general examination is usually administered in the January general examination period of the second year of study, after students have met all formal course requirements. This three-hour oral examination is administered by three faculty members from the graduate program, none of whom may be the student’s thesis adviser. The examination consists of two parts: the thesis proposal and second topic.

The thesis proposal probes depth of knowledge in the chosen research field and examines the ability of the student to justify and defend the goals, significance, and the experimental logic and methods of the proposed plan.

The second topic, or mini-proposal, is a two-page written document that uses an assigned research paper as the foundation for a research proposal. The student will propose a question and experiments to follow up on the results and/or conclusions in the assigned second topic paper.

Qualifying for the M.A.

The Master of Arts (M.A.) degree is normally an incidental degree on the way to full Ph.D. candidacy and is earned after successfully passing all parts of the general examination. It may also be awarded to students who, for various reasons, leave the Ph.D. program, provided that the following requirements are met:  completion of the formal courses and three laboratory rotations required for Ph.D. students, and demonstration of an appropriate level of research competency. Research experience must include at least one year of independent work in the laboratory, and competency must be demonstrated in writing. A faculty mentor and the graduate committee must approve the master’s paper.

Students are normally required to teach in two undergraduate-level courses. The first assignment is normally a laboratory course, while the second is normally a major undergraduate lecture course. Students may have the opportunity to do additional teaching if they wish to gain more experience.

Post-Generals requirements

Committee Meetings Each graduate student chooses a thesis committee that consists of the thesis adviser and two other faculty members who are knowledgeable in the student’s area of research. The thesis committee meets formally with the graduate student at least once per year, and sometimes more frequently on an informal basis. The responsibility of this committee is to advise students during the course of their research.

Dissertation and FPO

When the research is completed, the student writes the dissertation, which is first read by the adviser then by two additional readers chosen by the student. Usually, the second readers are the other members of the student’s thesis committee. Upon acceptance of the dissertation, the student gives a final, public oral presentation of the research to the department.

The Ph.D. is awarded after the candidate’s doctoral dissertation has been accepted and the final public oral examination sustained.

Program Offering: M.D./Ph.D.

Program description.

The Princeton Graduate School has a partnership with the Robert Wood Johnson Medical School (RWJMS) and the Rutgers University (New Brunswick) Graduate School of Biomedical Sciences to serve as a Ph.D. site for students enrolled in the M.D./Ph.D. program of RWJMS.

Students admitted to the M.D./Ph.D. program at RWJMS perform laboratory rotations at Princeton during the summer before and the summer after the first year of the pre-clinical portion of the program, prior to their enrollment as doctoral students, and subject to the approval of the faculty member and both the Princeton and the Rutgers MD/PhD program co-directors. Following the second rotation, a student will choose a laboratory for Ph.D. research by mutual agreement with a faculty adviser and approval by the Princeton MD/PhD program co-director, the chair of the Molecular Biology admissions committee, and the Graduate School.

Students who are accepted to work with a faculty member in, or an affiliated faculty member of, the Department of Molecular Biology will enter the Ph.D. program and receive that degree from Princeton. These students will fulfill Graduate School and departmental requirements, including the one-year residence requirement, taking and passing the general examination, and sustaining the final public oral examination. (Pre-clinical coursework at RWJMS will typically substitute for the department’s core curriculum.)

The Ph.D. portion of the joint program is expected to take three to four years. Extension beyond a fourth year requires approval from the Academic Affairs Committee of the joint degree program.

For those students considering the dual degree program, please take time to review the M.D./Ph.D. General information page .

M.D./Ph.D. students in the Department of Molecular Biology must take two courses, which can be either core or elective courses, achieving an average of B or better. Students may take additional elective courses that are closely related to their research topic.

Molecular Biology Annual Retreat The molecular biology annual retreat is a two-day symposium of research talks and poster sessions held in the fall and attended by all graduate students, postdoctoral fellows, and faculty in molecular biology and associated departments.  

The general examination is usually administered in the September/October general examination period of the second year of study, after students have met all formal course requirements. This two-hour oral examination is administered by three faculty members from the graduate program, none of whom may be the student’s thesis adviser.

The examination consists of a thesis proposal which probes the depth of knowledge in the chosen research field and examines the ability of the student to justify and defend the goals, significance, and the experimental logic and methods of the proposed plan.

The Master of Arts (M.A.) degree is normally an incidental degree on the way to full Ph.D. candidacy and is earned after successfully passing all parts of the general examination. It may also be awarded to students who, for various reasons, leave the Ph.D. program, provided that the following requirements are met: completion of the formal course requirements and demonstration of an appropriate level of research competency. Research experience must include at least one year of independent work in the laboratory, and competency must be demonstrated in writing. A faculty mentor and the graduate committee must approve the master’s paper.

When the research is completed, the student writes the dissertation, which is first read by the adviser then by two additional readers chosen by the student. Usually the second readers are the other members of the student’s thesis committee. Upon acceptance of the dissertation, the student gives a final, public oral presentation of the research to the department.

• Bonnie L. Bassler

Associate Chair

• Jean E. Schwarzbauer

Director of Graduate Studies

• Ileana M. Cristea
• Danelle Devenport (acting) (fall)

Director of Undergraduate Studies

• Elizabeth R. Gavis
• Rebecca D. Burdine
• Danelle Devenport
• Zemer Gitai
• Frederick M. Hughson
• Martin C. Jonikas
• Michael S. Levine
• Lydia Lynch
• Coleen T. Murphy
• Alexander Ploss
• Paul D. Schedl
• Stanislav Y. Shvartsman
• Thomas J. Silhavy
• Jeffry B. Stock
• Ned S. Wingreen

Associate Professor

• Mohamed S. Abou Donia
• Sabine Petry
• Jared E. Toettcher
• Martin Helmut Wühr

Assistant Professor

• Brittany Adamson
• John F. Brooks
• Michelle M. Chan
• Ricardo Mallarino
• Cameron A. Myhrvold
• Eszter Posfai
• AJ te Velthuis

Associated Faculty

• José L. Avalos, Chemical and Biological Eng
• Lisa M. Boulanger, Princeton Neuroscience Inst
• Clifford P. Brangwynne, Chemical and Biological Eng
• Mark P. Brynildsen, Chemical and Biological Eng
• Daniel J. Cohen, Mechanical & Aerospace Eng
• Jonathan M. Conway, Chemical and Biological Eng
• Thomas Gregor, Physics
• Ralph E. Kleiner, Chemistry
• A. James Link, Chemical and Biological Eng
• Lindy McBride, Ecology & Evolutionary Biology
• Tom Muir, Chemistry
• Celeste M. Nelson, Chemical and Biological Eng
• Joshua D. Rabinowitz, Chemistry
• Mohammad R. Seyedsayamdost, Chemistry
• Joshua W. Shaevitz, Physics
• Stanislav Y. Shvartsman, Chemical and Biological Eng
• Mona Singh, Computer Science
• Howard A. Stone, Mechanical & Aerospace Eng
• John D. Storey, Integrative Genomics
• Olga G. Troyanskaya, Computer Science
• Samuel S. Wang, Princeton Neuroscience Inst
• Bridgett M. vonHoldt, Ecology & Evolutionary Biology

Professor Emeritus (teaching)

• Sarah J. Flint

University Lecturer

• Heather A. Thieringer

Senior Lecturer

• Laurel Lorenz
• Jodi Schottenfeld-Roames
• Anthar S. Darwish
• Karin R. McDonald

Visiting Lecturer with Rank of Professor

• John J. Tyson

For a full list of faculty members and fellows please visit the department or program website.

Permanent Courses

Courses listed below are graduate-level courses that have been approved by the program’s faculty as well as the Curriculum Subcommittee of the Faculty Committee on the Graduate School as permanent course offerings. Permanent courses may be offered by the department or program on an ongoing basis, depending on curricular needs, scheduling requirements, and student interest. Not listed below are undergraduate courses and one-time-only graduate courses, which may be found for a specific term through the Registrar’s website. Also not listed are graduate-level independent reading and research courses, which may be approved by the Graduate School for individual students.

CHM 542 - Principles of Macromolecular Structure: Protein Folding, Structure and Design (also MOL 542)

Cos 551 - introduction to genomics and computational molecular biology (also mol 551/qcb 551), mat 586 - computational methods in cryo-electron microscopy (also apc 511/mol 511/qcb 513), mol 504 - cellular biochemistry, mol 506 - cell biology and development, mol 514 - molecular biology, mol 518 - quantitative methods in cell and molecular biology, mol 523 - molecular basis of cancer, mol 540 - research projects in molecular biology (laboratory rotations), mol 541 - research projects in molecular biology (laboratory rotations), mol 559 - viruses: strategy and tactics, mol 561 - scientific integrity in the practice of molecular biology, mol 567 - electron microscopy in structural biology (half-term) (also mse 542), neu 501a - cellular and circuits neuroscience (also mol 501a), neu 501b - neuroscience: from molecules to systems to behavior (also mol 501b), neu 502a - systems and cognitive neuroscience (also mol 502a/psy 502a), neu 502b - from molecules to systems to behavior (also mol 502b), neu 503 - neurogenetics of behavior (also mol 503), neu 537 - systems neuroscience: computing with populations of neurons (also mol 537/psy 517), qcb 515 - method and logic in quantitative biology (also chm 517/eeb 517/mol 515/phy 570).

Boston University Academics

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• PhD in Molecular Biology, Cell Biology & Biochemistry

The PhD in Molecular Biology, Cell Biology & Biochemistry (MCBB) is designed to build a solid foundation in related fields in the molecular biosciences through coursework and seminars, and to develop the skills for achieving cutting-edge research accomplishments. The program is based on an interdisciplinary environment and a breadth of approaches and biological interests, including faculty from several departments (e.g., Biology, Biomedical Engineering, Chemistry, Health Sciences). The primary goal of the degree is to prepare students for successful careers in academia, research institutions, or industry (e.g., biotechnology and pharmaceutical industries), or in government as policymakers. The program accepts applicants with degrees in related disciplines where the focus has been on biology, chemistry, biotechnology, biochemistry, or a relevant field.

Learning Outcomes

• Demonstrate academic mastery in the interrelated biological disciplines encompassing at least two of three biological areas: 1) molecular biology, 2) cell biology, and 3) biochemistry, plus in computational analysis.
• Achieve research expertise and complete a body of original research that advances a specific field of study in the molecular biosciences.
• Follow the ethics for appropriate behavior in the scientific discipline.
• Be able to teach and promote the field of biological sciences.
• Be prepared to enter the job market.

Course Requirements

For details on requirements regarding grade point averages, annual reports, and responsible conduct in research, please see the MCBB graduate program guide .

A total of 64 units is required. Of these, a minimum of 32 units must derive from lecture or seminar courses, and a minimum of 12 units from elective courses. Remaining coursework normally consists of research units. Incoming students with previous graduate units, including from a master’s degree at a different institution, may apply to transfer course units toward the PhD degree. Course requirements are as follows:

• CAS BI 735 Advanced Cell Biology
• CAS BI 753 Advanced Molecular Biology
• CAS MB 583 Progress in Cell & Molecular Biology (first term)
• CAS MB 584 Progress in Cell & Molecular Biology (second term)
• CAS MB 697 A Bridge to Knowledge: A Practical Seminar for First-Year Graduate Students
• CAS MB 721 Graduate Biochemistry or MB 722 Advanced Biochemistry
• A graduate-level quantitative course from a list of options
• 12 units in elective courses

Language Requirement

There is no foreign language requirement for this degree.

Qualifying Examination

This examination is designed to test the student’s general knowledge encompassing molecular biology, cell biology, and biochemistry, as well as the student’s grasp of the proposed research project and ability to synthesize findings and propose and experimentally test hypotheses. It consists of an oral qualifying exam where the student will defend a written research proposal. The exam is administered by a committee of no fewer than five faculty, at least four of whom must be members of the MCBB program (including the thesis advisor). The qualifying exam should be completed within 2.5 years of matriculation into the MCBB PhD program. Any student failing this examination, at the discretion of the examination committee, may have the opportunity to take it again. Failure of the second examination will be grounds for automatic dismissal from the PhD program and the loss of any further financial aid, although the student may still be eligible for the MA provided that those degree requirements are met.

Dissertation and Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation that presents original research or creative scholarship. A prospectus for the dissertation must be approved by the readers, the Director of Graduate Studies, and the Department Chair/Program Director. Candidates must pass a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the GRS General Requirements for the Doctor of Philosophy Degree .

Teaching Requirement

The PhD program requires a minimum of two terms of teaching during a graduate student’s career.

Students electing to leave the PhD program may leave with a master’s degree if they have completed their core course requirements, have obtained 32 units, and have passed their Qualifying Exam, or have otherwise satisfied the requirements of the master’s degree .

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Department of Molecular and Cell Biology

Genetics and Genomics

Formal “qualification” for the Ph.D. degree takes place by passing the Dissertation Proposal, a tripartite examination focused upon the student’s dissertation research plans. This exam should be taken at a point at which the student has completed most course work and has research well underway. The student should aim to complete this exam by the end of the third year of graduate study. The three parts of the exam, each of which will be evaluated separately by the full Advisory Committee are:

I. A written proposal II. A seminar presentation on the proposal III. A closed-door question and answer session with faculty

A student who demonstrates acceptable performance on all three parts of the examination, evidenced by a majority vote of the full Advisory Committee to pass on all three sections, “qualifies” for the Ph.D. degree, and continues on that track of study. A student who does not make adequate progress, evaluated by a majority vote of the Committee, may be asked by the Committee to repeat any sections of the examination to achieve a full pass. In cases of inadequate performance on the examination, the Committee may also recommend transfer to one of the Master’s of Science programs.

I. GUIDELINES FOR WRITTEN PROPOSAL PREPARATION

(For Genetics and Genomics doctoral scholars in MCB)

The written proposal has a ten page limit ( excluding references) and the following suggested sections. All figures, tables, charts, and diagrams are included in this 10-page limit. This format is based on current grant submission formats for most federal agencies, which range from 4-12 pages total, preparing the student for succinct presentation and defense of their scientific premise.

You must submit this Proposal two calendar weeks (10 business days) before the scheduled examination to each of your committee members. The thesis advisor may read and make general comments on this document prior to submission, but may not edit it. For some guidelines on writing, Helpful Hints on Scientific Writing.

Cover page: This is not included within the 10-page limitation.

It will include: Title, date of submission, date of scheduled exam, student name, committee members’ names and affiliations.

I. Significance. What are the broad implications of the research that you propose? What is its importance? The significance section should “funnel” consideration from the global to the specific project at hand. One warning: everything you mention in this section is fair game for questioning. Keep focused on the issues you identify as really important. (1/2 – one page)

II. Specific Aims/Goals. Make use of numbered, concise statements of hypotheses/questions. This will immediately focus the reader on precisely what you will be doing, and place the background in context. Keep in mind that this does not have to reflect historical chronology, but rather should present a series of logical steps. (1/2 -one page)

Sections I-II is the total content of Page 1 and cannot exceed one page.

Pages 2-10 Consist of the Following Sections:

III. Background and Preliminary Data. Provide a brief synopsis of the relevant background the reader needs to interpret your proposed research. (2 pages or less) This should not be an exhaustive literature review, but rather should highlight the background needed to place the area of research into context to understand your experimental hypotheses and approaches.

Keep in mind not all members of your committee are in the same area of research; it is critical to explain why the system/question/approaches proposed are interesting, important, and feasible.

In the preliminary data component of this section, a brief presentation of the data collected by the student in support of the approach and aims should be included. Note that considerable variation in the extent of data among students is expected, but only include data relevant to the proposal.

IV. Approach. This section is the bulk of the proposal (4-5 pages). It is a good idea to have a subsection for each hypothesis/question posed in each specific aim. In this section, you are tasked with defending why you should continue for the next 2-4 years on your project. In other words, convince your readers that this work is worthwhile, feasible and will contribute to the field.

Include the following subsections under each aim in the approach section:

A. Rationale . This is a statement of the logic behind your experiment. Include in this section any thinking that went into your hypothesis, any synthesis you might have made.

B. Experimental Plan. Include in this section the strategy you plan to use to address the hypothesis, as well as information about procedures and protocols in general terms. Your committee is more interested in the logic than in the details – reference common procedures. Focus on those aspects that are conceptual rather than technical, but be aware of any limitations of the methodology you select.

C. Interpretations and Alternative Approaches. Make sure you interpret results critically. Showing alternative meanings indicate that you have thought the problem through and are able to meet future challenges. Call attention to potential difficulties you may encounter with each approach. Propose alternatives that would circumvent possible limitations. Committee members will be aware of possible problems; convince them you can handle such circumstances.

For example:

Specific Aim 1: To…

1.A. Rationale – why do this? 1.B. Experimental Plan – how will I do this? 1.C. Interpretations and alternative approaches – what will it mean if I see X or Y? If it does not work because of the following reason…I will perform….to overcome this problem

IV. Timeline and Impact.

In this section, briefly lay out your timeline of experiments for the remainder of your thesis, including anticipated milestones such as publication submissions, conference presentations, and other seminar opportunities. Do not include courses, teaching and other duties not directly relevant to the work.

The impact statement should summarize (2-4 sentences total) what your body of work would contribute to the field, highlighting the advances it makes over existing knowledge.

II. SEMINAR PRESENTATION of PROPOSAL (see Presentation Skills )

Iii. closed door exam.

Molecular and Cellular Biology

Share this page.

In this interdisciplinary program, you will interact with students and faculty who have diverse backgrounds in chemistry, marine biology, computational biology, and others. The skills you will learn in your first year include coding, how to write fellowships, and how to give scientific talks to a non-science audience. You will begin bonding with your small cohort of fellow students with an annual trip to Cape Cod to talk about science at the beginning of the academic year.

Dissertations that students have completed include "Characterization of the Epigenetic Regulator LSD1 as a Druggable Dependency in Treatment of Resistant Melanoma" and "Circadian Clocks in the Real World: Effects of Dynamic Light Regimes on the Regulation of Circadian Gene Expression in Cyanobacteria."

Graduates have secured positions in industry with companies such as Google, Genentech, and Novartis. Others serve in academic positions at many prestigious schools including Harvard, Caltech, Stanford, and MIT. 

Additional information on the graduate program is available from the Department of Molecular and Cellular Biology , and requirements for the degree are detailed in Policies .

Areas of Study

Molecules, Cells, and Organisms

Admissions Requirements

Please review the admissions requirements and other information before applying. You can find degree program-specific admissions requirements below and access additional guidance on applying from the Department of Molecular and Cellular Biology .

Academic Background

Applicants should have a record of introductory courses in chemistry, biology, physics, and mathematics. While the following courses should not be regarded as prerequisites for admission to graduate study, most admitted students have completed these courses as undergraduates:

• Biology (at least one general course in biology and two terms of biology at a more advanced level)
• Biochemistry
• Organic chemistry
• Physical chemistry
• Physics (a general course)
• Mathematics (a basic knowledge of differential and integral calculus). Competence in elementary programming is also desirable.
• Laboratory in biology, biochemistry, or instrumental analysis.

Statement of Purpose

Describe your reasons and motivations for pursuing a graduate degree in your chosen degree program, noting the experiences that shaped your research ambitions, indicating briefly your career objectives and concisely stating your past work in your intended field of study and in related fields. Your statement should not exceed 1,000 words.

In addition to the above guidance, your statement of purpose should also address the following questions:

• The focus of this question should be a discovery in which you had substantial engagement and personal impact on the research. Do not reference a large group project simply because it was interesting. 
• If you have not had significant research experience, please describe a scientific discovery that motivated you to pursue research. 
• Using simple language, describe what you or others did, why, and what it means.
• Molecular Mechanism
• Cell and Developmental Biology
• Molecular Ecology and Evolution
• Choose two MCO faculty members that you are interested to work with and explain why by using a specific example from their published work.

Personal Statement

Standardized tests.

GRE General: Not Accepted iBT TOEFL preferred minimum score: 100 IELTS preferred minimum score: 7

Theses and Dissertations

Theses & Dissertations for Molecular and Cellular Biology

See list of Molecular and Cellular Biology faculty

APPLICATION DEADLINE

Questions about the program.

Department of Molecular Biosciences, The Wenner-Gren Institute

Phd studies at mbw.

Welcome to the internal pages for PhD studies at MBW. The information on this page is aimed at PhD students and supervisors, and covers everything from credit registration to preparation of the public defence.

Instructions for group leaders on how to employ a PhD student, can be found here

On this page you will find useful information, templates and checklists regarding PhD studies at MBW. 

PhD Council and PhD Council Board

The PhD Council at the department of Molecular Biosciences, The Wenner-Gren Institute (MBW) represents about 60 graduate students covering three different profile areas: Infection and Immunobiology, Integrative Biology and Molecular Cell Biology. The PhD Council’s primary function is to ensure quality of education and to promote good working and social environment for our graduate students.

The PhD Council Board is responsible for the overall operations of the PhD Council. The board meets regularly and consists out of seven graduate students:

• Max Louski (Chair)
• Isabella Badolati (Vice Chair)
• Simon Prokisch (Secretary/Treasurer)
• Tajana Lömker (Board member)
• Sergei Pirogov (Board member)
• Anaswara Sugathan (Board member)

If you want to get in contact with the PhD Board, please send an e-mail to [email protected] .

Protocols PhD Council and PhD Council Board

Application for PhD studies

General Syllabus

The subject Molecular Biosciences at Stockholm University includes studies and research in three transdisciplinary profile areas: 1) molecular cell biology, 2) infection and immunobiology, and 3) integrative biology. The general syllabus regulates the PhD program at MBW. In case of discrepancy between the Swedish original and the English translation of the syllabus, the Swedish version takes precedence. 

PhD Guide - to new graduate students

The PhD Guide reflects a combined MBW/PhD Council effort and aims to help new graduate students to navigate through the various milestones that lead to a PhD degree. 

Individual study plan (ISP)

In accordance with the Higher Education ordnicance, each doctoral student must have an Individual study plan (ISP). The ISP is an agreement between the PhD student and the supervisor, and regulates the content of the research and studies.

The first Individual study plan 

The first Individual study plan is drawn up prior to admission, and is formally decided upon after the admission. 

Yearly follow-up of the ISP

The ISP is updated annually in conjunction with a seminar given by the student. The members of the ISP follow-up committee should attend the seminar. The first update should take place approximately six months after the start of the program. The ISP-form and the protocol is handed in to the administration after the follow-up. 

Obligatory courses at MBW

There are three obligatory courses for PhD Students at MBW. 

• Introduction to research studies in Biology

Registration of credits for PhD students

You get credits by attending courses, seminars or conferenes, giving a talk or presenting a poster. Your supervisor needs to approve these activities and you need to report them to Head of Research Subject, Kristina Jonas, in order for them to be registered in Ladok (the study administrative system). You need at least 50 credits in order to be able to procced to your defence. 

Travel grants

Travel grants - stockholm university .

Grant application are ranked by the Head of the department and assessed by the stipend committee. There is usually no need for long essays or multi-page CVs, but make sure to include the most important information, also about the course or meeting you want to attend. The final results are usually announced a few month after the respective deadline.

MBW Travel grant

Read more about travel booking, accomodation etc

Prolongation of study time

In Sweden, PhD students are financed for 48 months of full-time studies/thesis research. At MBW, the PhD studies are financied for 5 years (60 months full-time) with the costs distributed on the basis of 80% full-time studies/thesis research and 20% assistantship, including teaching. Read more about prolongation of the study time in the documents below. 

Half time review and licentiate

When the PhD student has carried out approximatley two years of full-time doctoral studies, a half-time review or a licentiate seminar should take place. At MBW, the half-time review is the most common option. 

Half time review

The aim of the half-time review is to assess the general feasibility of the projects leading to a PhD, the student’s general knowledge in the subject of research, the knowledge specific to the project, and the ability of the student to present research results in written and oral forms.

When the half-time review is completed, the form “Half-time protocol” is filled in by the committee. The protocol is sent to the head of department/the department’s director of doctoral education and is attached to the doctoral student’s file. A copy of the “Half-time protocol” is sent to the doctoral student and supervisors.

Licentiate seminar

The instructions for the licentiate examination are in accordance with the General Syllabus and follow the guidlines of the Faculty of Science. See the instructions below: 

Read more about the rules concerning the licenciate examination (in Swedish)

Dissertation

The public defence is an examination and a highlight to mark the end of four years of full-time study. In the sections below you will find forms and checklists to help you prepare for your defence. 

Regulations - public defence of a doctoral thesis

In order to ensure that the review of the thesis and the public defence seminar is impartial, objective, and academically sound, the Board of Science has adopted the following regulations in addition to the regulations of the Higher Education Ordinance and the University.

Regulations for public defence of doctoral thesis in the disciplinary domain of Science

Checklist and timeline

The checklist should be seen as a complement to the Regulations for public defence, and describes the practical routines and division of tasks at MBW. Read this checklist througouly well in time before the defense. The checklist contains links to several of the forms that are also listed below (for example "Public defence form" and "Review and recommendation"). 

Printing support

The University Library supports the process of getting theses and dissertations ready.

Read more and view the timeline for printing support

Review and recommendation

As described in point 1 in the checklist, the review of achievements and material to be included in the thesis is the first step to take when approaching your defence. The ISP follow-up committee shall review the achievements of the PhD student and the papers that are to be included in the doctoral thesis with the aim of issuing a recommendation to proceed or not with the thesis defence. 

Public defence form

As described in point 3 in the checklist, the supervisor should fill in the application form for public defence, and hand it to the head of the department at least seven weeks prior to the defence. Make sure that all the prior points in the checklist have been ticked off before you proceed with this. 

Public defence form ("Förslag på opponent, betygsnämnd, ordförande och kontaktperson för disputation inom det Naturvetenskapliga området")

Information to opponents

As described in point 4 in the checklist, information to the opponent may be sent prior the defence. Below you will find two documents with general guidlines on the dissertation act, as well as practical information concerning transportation and accommodation for example. 

Pre-review of the thesis

As described in point 13 in the checklist, the opponent and the members of the examining board must receive a copy of the thesis at least two weeks before the thesis is printed. The template letter below can be used for this purpose. 

Apply for your diploma

After a succesful defence, you will be awarded a degree certificate upon request.

Please follow the instructions provided here to apply for your degree certificate

Last updated: August 15, 2024

Source: MBW

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Biochemistry, Cellular and Molecular Biology, PhD

School of medicine, fields of study and research.

The Departments of Biological Chemistry, Biophysics and Biophysical Chemistry, Cell Biology, Molecular Biology and Genetics, Neuroscience, Pharmacology and Molecular Sciences, and Physiology jointly provide a program of study leading to the PhD degree. The objective of the Biochemistry, Cellular and Molecular Biology (BCMB) program is to provide students with the breadth of knowledge and research training that will prepare them for their own independent and productive careers.  Strong emphasis is placed on molecular and cellular approaches to fundamental problems in research areas covered by the participating departments. The focus on a broad background in basic science for all students promotes the study of problems of biomedical importance from a mechanistic perspective. The multi-departmental nature of the program fosters a collaborative environment that is ideal for graduate training, and promotes the extensive interaction and collegiality that is a hallmark of Johns Hopkins science.

Training includes a rigorous and broad-based first year curriculum, research rotations in three different laboratories, a proposal-based qualifying exam (along with a proposal-writing workshop), and thesis research in the laboratory of one of more than 100 participating faculty members. Once they join a laboratory, students establish a departmental affiliation and participate in their departmental journal clubs and seminars. At the same time, students continue to maintain their BCMB identity and affiliation in a number of ways. These include attending the annual BCMB retreat, assisting in the recruitment of new students, hosting a one-day symposium with outside speakers, and participating in a monthly student research colloquium (organized by BCMB students for their peers). There is no formal teaching requirement for BCMB students, but many opportunities for gaining teaching experience are available.

Due to the multidisciplinary focus of the BCMB program, students are very well prepared for the full breadth of modern biomedical science careers. Greater than 90% of students in the BCMB program complete the PhD degree. The average time to degree over the last 10 years is 5.7 years. On average, BCMB students publish 4 research papers with 2 as first author.

To date, there are over 750 BCMB graduates; many hold leadership positions in academia, industry, government and other sectors. At least 97% of graduates in the last 5 years are following career paths in laboratory research, medicine, or research-related careers.

The lecture halls and the research laboratories are located in the Wood Basic Science Building and the adjoining Physiology, Biophysics, Hunterian, Preclinical Teaching, Ross Building, Miller Research, and Rangos Buildings. Modern equipment and state-of-the-art core facilities are available to students. This includes electron microscopes, confocal microscopes, nuclear magnetic resonance spectrometers, mass spectrometers, HPLC and gas-liquid chromatographs, protein sequencers, peptide synthesizers, oligonucleotide synthesizers, X-ray diffraction equipment, as well as instrumentation for microarray analysis and deep sequencing.

Collaborative Facilities

All seven departments enjoy excellent working relationships with other departments in the medical school and with departments at the Homewood campus.

Financial Support

All students accepted into the BCMB program receive full tuition, health and dental insurance, and stipend support for the duration of their studies.

Admission Requirements

Candidates for admission should show a strong academic foundation with coursework in biology, chemistry, physical sciences, and quantitative analysis. A bachelor's degree from a qualified college or university will be required for matriculation.

The BCMB website has up-to-date information on “ How to Apply ” and “ Frequently Asked Questions .” For questions not addressed on these pages, please email [email protected] .

Program Requirements

Students must successfully complete all eight courses of the “Foundations of Modern Biology” core curriculum, and the Core Discussion course in the first three quarters of year one. During the fourth quarter, students complete two electives, the “BCMB Responsible Conduct in Research” course, and the “Method, Logic and Experimental Design” course.  Three additional elective courses are required in the advanced years of study in addition to the BCMB "Seminar" and "Writing a Paper" courses.

An oral examination, conducted as prescribed by the Doctor of Philosophy Board of the University, must be completed before the end of the second year. Subsequently, students are required to participate in annual thesis committee meetings as they work on their thesis project, to review research progress and discuss plans for the next year.​​​

Usually in year five, the student's thesis committee agrees that the student is nearing completion of their research and will be ready to write a dissertation. The student's advisor and one other member from their thesis committee will read and approve the dissertation. The student must present a public seminar on their completed thesis research.

The BCMB program recognizes the importance of students’ professional development and career exploration during their graduate studies. In order to better integrate professional development into the training, the BCMB program is formally participating in the “OPTIONS” Career Curriculum offered by the Professional Development and Career Office (PDCO). During the first two years, students attend workshops and panel discussions facilitated by professionals in different fields discussing their career paths, what each profession entails, how the graduate training relates to the required skills and knowledge, and what additional skills would be useful. Several of the speakers are BCMB program alumni, and offer to follow-up with interested students, fostering a sense of community. In year 3, students join an OPTIONS Career Community which comprises six monthly workshops that provide exposure to careers and skills assessment (e.g., Academic Research; Business and Finance; Biotech and Pharma; Science Communication and Policy; Academic Teaching; etc.). In year 4 and beyond, students meet with a career coach to create a plan for gaining more career-relevant experiences and/or participate in an internship.

The first year required courses are as follows:

Courses are detailed under the course descriptions listed in the entries of the Departments of Biological Chemistry, Biophysics and Biophysical Chemistry, Cell Biology, Molecular Biology and Genetics, Neuroscience, Pharmacology and Molecular Sciences, and Physiology, or as Interdepartmental.

Microbiology and Molecular Biology

Application deadline.

• Fall: Jan 15

Contact Information

• Statement of intent explicitly stating field of interest and career goals
• Baccalaureate degree or equivalent in microbiology or biological or biochemical sciences
• Classes in chemistry, physics, mathematics, and statistics are recommended
• Employment History
• Curriculum Vitae (CV)
• Cover Letter
• Copies of Publications
• List of Presentations
• Foreign credential evaluation for international students
• English proficiency is required for most foreign applicants. We will not consider applicants whose English proficiency is below the minimum scores required by the university.
• PhD after BS: 54 credit-hours (at least 18 didactic credit hours, 18-20 research credit-hours, and 18 dissertation credit hours [MMBIO  799R ]).
• PhD after BYU MMBio MS: 36 credit-hours (18-20 research credit hours and 18 dissertation credit hours [MMBIO  799R ]).
• Cell 570 - Responsible Conduct of Research, 1.0
• MMBIO 660 - Microbiology and Immunology, 3.0
• MMBIO 661 - Molecular Genetics in Practice, 2.0
• MMBIO 663 - Articulating Science, 2.0
• MMBIO 665 - Genomics, 3.0
• MMBIO 691R  - Graduate Seminar, 2.0
• MMBIO 692R - Research in Progress, 2.0
• MMBIO 695R - Research, 18.0 - 20.0
• MMBIO  799R - Doctoral Dissertation, 18.0
• Bio  503 - Research orientation, 1.0
• MMBIO  510  - History and Philosophy of MMBIO, 2.0
• MMBIO  518 - Select Pathogens, 2.0
• MMBIO  522 - Flow Cytometry, 2.0
• MMBIO  528R - Scientific Literacy, 1.0
• MMBIO  551R - Current Topics in MMBio, 1.0-3.0
• MMBIO 667 - Quantitative Genomics, 2.0
• Semiannual progress reviews.
• Presentation of research 1-2 times per year during Research in Progress
• Qualifying Examination: Student will be required to pass the qualifying exam. This includes (1) written outside grant proposal outside the area of expertise, and (2) oral exam on coursework and grant defense.
• Dissertation: standard university dissertation.
• Public seminar and oral defense of dissertation.

Department Courses Department Faculty

We have 69 Molecular Biology (proposal) PhD Projects, Programmes & Scholarships

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Molecular Biology (proposal) PhD Projects, Programmes & Scholarships

Nihr newcastle biomedical research centre and haleon phd studentship: investigation into the links between periodontal disease and inflammatory bowel disease, phd research project.

PhD Research Projects are advertised opportunities to examine a pre-defined topic or answer a stated research question. Some projects may also provide scope for you to propose your own ideas and approaches.

Funded PhD Project (Students Worldwide)

This project has funding attached, subject to eligibility criteria. Applications for the project are welcome from all suitably qualified candidates, but its funding may be restricted to a limited set of nationalities. You should check the project and department details for more information.

Deciphering the non-homologous end joining (NHEJ) DNA-double strand break repair mechanism in Mycobacterium tuberculosis

Self-funded phd students only.

This project does not have funding attached. You will need to have your own means of paying fees and living costs and / or seek separate funding from student finance, charities or trusts.

Understanding γδ T cell response in obesity-associated hepatocellular carcinoma (HCC)

Redefining dna-double strand break repair in humans, molecular mechanisms and targeted therapy for spinocerebellar ataxia [self-funded opportunity], join our graduate school at the interface of biology and computation: imprs-bac in berlin, funded phd programme (students worldwide).

Some or all of the PhD opportunities in this programme have funding attached. Applications for this programme are welcome from suitably qualified candidates worldwide. Funding may only be available to a limited set of nationalities and you should read the full programme details for further information.

Max Planck Research Programme

Max Planck Research Programmes are structured PhD opportunities set up by the Max Planck Society, an independent non-profit German research organisation. Max Planck Institutes and universities collaborate to offer interdisciplinary and international PhD opportunities providing high standards of training and support as well as generous funding.

Genetics of interstitial lung disease

Obesity, type 2 diabetes and the thrifty genotype hypothesis, identifying the causes and cellular effects of focal segmental glomerulosclerosis (fsgs), understanding and curing genetic kidney disease, assessing neuroinflammation and vascular dementia in association with atherosclerosis and aneurysm risk, exploring the contribution of divergent neuroinflammation in alzheimer’s disease, does the brain renin angiotensin system influence the occurrence of depression in alzheimer’s disease, investigating associations between the brain renin angiotensin system and traumatic brain injury, studies of brain renin angiotensin system changes in parkinson’s disease to find new treatments.

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