phd student biologie

Graduate Admissions

Helpful guides for applying to the biology phd program, tips for applying to the cmob track, tips for applying to the eco/evo track.

The PhD program trains students for careers in research and teaching. In addition to a wide range of courses, the PhD program offers informal but intensive instruction in laboratories and seminars, characterized by close contact between students and their faculty advisors. Biology is one of fourteen home programs  in the  Biosciences,  which span the School of Medicine and the School of Humanities and Sciences.

How to Apply

For the 2023-2024 PhD application season, application processes will be run through existing Biology (School of Humanities & Sciences)  and the Oceans Department(Stanford Doerr School of Sustainability). 

Previously part of the Biology Department , Hopkins Marine Station is now part of the Oceans Department within the Stanford Doerr School of Sustainability (SDSS).  Several faculty took appointments in Oceans, while others chose to stay in Biology; still others have joint appointments.  Additionally, several faculty members in Ecology/Evolutionary Biology now have joint appointments within Biology and SDSS. Based on these changes, the following information will provide further guidance for the application process. 

• Faculty with 100% appointments in Biology, will only recruit through Biology.  The Biology application will be used.
• Ecology/Evolutionary Biology faculty with 50/50 appointments in both Biology and SDSS will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Ecology/Evolutionary faculty with joint Biology/SDSS appointments, you should reach out to them directly regarding the appropriate application track to choose. 
• 100% Oceans faculty will only be recruiting students through the Earth Systems Science application.
• Oceans faculty appointed 50/50 in Biology and SDSS (Oceans Department), will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Oceans faculty with joint Biology/SDSS appointments, you should reach out to the individual faculty members you hope to work with regarding the appropriate application and track to choose. 

Faculty Affiliated with the Stanford Doerr School of Sustainability

Oceans Department:

Fio Micheli 100% 

Jeremy Goldbogen 100%

Giulio De Leo 100%

Larry Crowder 100%

Steve Palumbi 50% Oceans/50%Biology

Barbara Block 50% Oceans/50% Biology

*Note Chris Lowe will remain 100% in Biology

Ecology/Evolution

Liz Hadly 50% SDSS/50%Biology

Rodolfo Dirzo 50% SDSS/50%Biology

Tad Fukami 50% SDSS/50%Biology

Kabir Peay 50% SDSS/50%Biology

Chris Field 50% SDSS/50% Biology

Gretchen Daily 50% SDSS/50% Biology

Application and Deadline

Stanford Biology PhD Program applications are made through Graduate Admissions . The application deadline for Autumn Quarter 2024 matriculation is December 4, 2024 at 11:59pm pst.  The application for the Autumn 2025 cohort will be available in September 2024.  Please review the Graduate Admissions website prior to starting your application. Applications are considered for admission in Autumn Quarter only. 

Application Status

To check your application status,  click here to Visit Your Status Page .  

Requirements

• U.S. Bachelor's degree or its foreign equivalent
• Stanford Graduate application
• Non-refundable application fee of $125 (or fee waiver found here ) 
• Statement of Purpose
• Three Letters of Recommendation (a maximum of four letters will be accepted)
• Transcripts - Unofficial transcripts from all universities and colleges you've attended for one year or more must be uploaded to the online application.  Graduate Admissions only requires admitted applicants who accept the offer of admission to submit official transcripts that shows their degree conferral. Please do not send or have sent any official transcripts to us at this time.
• TOEFL Test of English as a Foreign Language (required for non-native English speakers)
• A current Curriculum Vitae/Resume

Knight-Hennessy Scholars

Join dozens of  Stanford School of Humanities and Sciences students  who gain valuable leadership skills in a multidisciplinary, multicultural community as  Knight-Hennessy Scholars  (KHS). KHS admits up to 100 select applicants each year from across Stanford’s seven graduate schools, and delivers engaging experiences that prepare them to be visionary, courageous, and collaborative leaders ready to address complex global challenges. As a scholar, you join a distinguished cohort, participate in up to three years of leadership programming, and receive full funding for up to three years of your PhD studies at Stanford. Candidates of any country may apply. KHS applicants must have earned their first undergraduate degree within the last seven years, and must apply to both a Stanford graduate program and to KHS. Stanford PhD students may also apply to KHS during their first year of PhD enrollment. If you aspire to be a leader in your field, we invite you to apply. Learn more about  KHS admission .

Biology PhD Program General Information

Diversity, inclusion, equity, and belonging.

The Department of Biology recognizes that the Supreme Court issued a ruling in June 2023 about the consideration of certain types of demographic information as part of an admission review. All applications submitted during upcoming application cycles will be reviewed in conformance with that decision. The Department of Biology welcomes graduate applications from individuals with a broad range of life experiences, perspectives, and backgrounds who would contribute to our community of scholars. Review of applications is holistic and individualized, considering each applicant’s academic record and accomplishments, letters of recommendation, and admissions essays in order to understand how an applicant’s life experiences have shaped their past and potential contributions to their field.Please see the websites below for more resources.

• Diversity and Engagement in Biosciences PhD Programs
• Vice Provost for Graduate Education
• Biology Equity and Belonging
• Research Sites
• Faculty Profiles and Labs
• Inquiries about the PhD in Biology should be directed to  biologyadmissions [at] lists.stanford.edu (biologyadmissions[at]lists[dot]stanford[dot]edu)

Biology, PHD

On this page:, at a glance: program details.

• Location: Tempe campus
• Second Language Requirement: No

Program Description

Degree Awarded: PHD Biology

The PhD program in biology offers individualized courses of study tailored to students' interests that include laboratory, field and theoretical work. Flexibility in the program is achieved by requiring only one core class, which is a choice between two topics that cover the breadth of the research directions in this program.

Electives are also as flexible as possible and include areas outside of biology. The major goal is to provide students with the opportunity to create an adaptable and general degree program that allows both traditional and interdisciplinary approaches in any area of biology.

Program Faculty   PhD Students

• BIO 792 Research
• BIO 514 Statistical Models for Biology
• BIO 522 Populations: Evolutionary Ecology
• BIO 507 Advanced Evolutionary Medicine
• BIO 521 Landscape Ecology
• BIO 530 Scientific Teaching (required in first semester for students that will TA at any point in degree)
• BIO 533 K-12 STEM Education & Outreach
• BIO 539 Computing for Research
• BIO 541 SOLS Seminar Series
• BIO 542 SOLS Current Topics in the Life Sciences
• BIO 591 Society and Natural Resource Management
• BIO 591 Social-Ecological Systems & Adaptation
• BIO 591 Drylands in a Changing Earth
• BIO 591 Population Genetic Reading Group
• BIO 591 Communication for Scientists
• BIO 591 Genetics and Genomics of Behavior
• BIO 620 Research Prospectus Writing
• EVO 501 Current Topics in Evolutionary Biology
• EVO 601 Principles of Evolution
• EVO 610 Research Areas of Evolution

Application and admission information

How to apply.

Applications open September 1 for admission in Fall of the following year. The application deadline is December 1 . We accept applications for Fall semesters only. We cannot guarantee that applications received after the December 1 deadline will be considered for admission.

All applicants must apply by filling out ASU's Graduate Admissions application. All application materials must be submitted through the application or to Graduate Admissions directly. Please do not mail or email any documents to the School of Life Sciences. 

Required materials and information include the following:

• 1-2 page personal statement
• An up to date CV or resume
• The names of relevant SOLS faculty you have been in touch with who you might be interested in being supervised by
• Unofficial transcripts and English proficiency test scores (if applicable)
• The names and emails of at least 3 recommenders to write you letters of recommendation

Application review process and timeline

Following the December 1 deadline, faculty will begin reviewing applications. Applicants should monitor their My ASU priority tasks to ensure there are no missing materials in their application.

Faculty will decide which applicants they would like to invite to our Graduate Recruitment Weekends (GRWs), typically held in February. Applicants will hear from the School of Life Sciences in January if they are invited to participate in the GRWs.

Admission decisions will begin after the GRWs, and applicants typically receive final decisions by April 1.

Requirements

Minimum requirements for admission include the following:

• Cumulative GPA of at least 3.0 on a 4.0 scale
• There are other ways to demonstrate English proficiency beyond the tests, so please refer to ASU's English proficiency webpage to review how you might satisfy requirements

Desired qualifications typically seen in competitive candidates:

• Research experience and a letter of recommendation from a faculty research supervisor
• English proficiency scores that meet these teaching assistant language proficiency requirements

Please note that the GRE is not required.

Students offered admission to a PhD program in the School of Life Sciences will typically receive a funding offer as well. While individual funding offers may differ to some degree, they typically include teaching assistant and/or research assistant positions each semester (summer optional) for 5 years. These positions provide financial coverage through the following:

• A standard salary stipend paid biweekly
• Tuition remission covering enrollment in 6-18 credit hours for fall and spring semesters and 1-14 credit hours for summer semesters
• Health insurance coverage

To discover more, check out the ASU Graduate College's funding opportunities !

Degree Requirements

Curriculum plan options.

• 84 credit hours, a written comprehensive exam, an oral comprehensive exam, a prospectus and a dissertation

Required Core (3 or 4 credit hours) BIO 514 Statistical Models for Biology (4) or BIO 620 Research Prospectus Writing (3)

Electives (67 or 68 credit hours)

Other Requirements (1 credit hour) BIO 610 Introduction to Responsible Conduct of Research in Life Sciences (1)

Culminating Experience (12 credit hours) BIO 799 Dissertation (12)

Additional Curriculum Information The program is tailored to the needs of the individual student. The plan of study is developed by the student and a supervisory committee consisting of a major professor and three additional faculty members. The electives credit hours are comprised of coursework showing breadth in biology subdisciplines, seminars and research.

An oral comprehensive examination is required in order to advance to candidacy, and a formal defense of the final written dissertation is required prior to graduation.

Admission Requirements

Applicants must fulfill the requirements of both the Graduate College and The College of Liberal Arts and Sciences.

Applicants are eligible to apply to the program if they have earned a bachelor's or master's degree in biology or a related discipline from a regionally accredited institution.

Applicants must have a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in the last 60 hours of their first bachelor's degree program, or a minimum cumulative GPA of 3.00 (scale is 4.00 = "A") in an applicable master's degree program.

Applicants must submit the following:

• graduate admission application and application fee
• official transcripts
• academic record form
• personal statement
• curriculum vitae or resume
• three letters of recommendation
• proof of English proficiency

Additional Application Information An applicant whose native language is not English must provide proof of English proficiency regardless of their current residency.

Research experience is desired for entry into this program.

Next Steps to attend ASU

Learn about our programs, apply to a program, visit our campus, learning outcomes.

• Able to communicate the findings and originality of their research, both orally and in writing.
• Able to perform research that they have designed to address important and challenging questions in the field of biology.
• Able to review scholarly literature as they develop specialized expertise in their selected research area.

Career Opportunities

Those who have earned a doctorate in biology are prepared for academic careers at every level, from community colleges to research universities. Their skills and knowledge are also valuable for government careers with federal and state agencies responsible for wildlife management and conservation, and for careers in industry and nongovernmental organizations.

Career examples include:

• food, agriculture and health care scientists in academic, private and industrial labs
• principal investigators in government labs and nonprofit organizations
• professors or instructors in universities and colleges
• science teachers in elementary and high schools
• wildlife, animal and conservation scientists

Program Contact Information

If you have questions related to admission, please click here to request information and an admission specialist will reach out to you directly. For questions regarding faculty or courses, please use the contact information below.

• [email protected]
• 480/965-1768

Ph.D. Program in Biology

Our graduate program provides students of diverse backgrounds with the opportunity to intensely engage in research in the biological sciences and contribute to the broader scientific community.

Average time to degree: 5.3 years

Percentage of graduates in post-doctoral or permanent positions: 94%

Average publications after completion of program: 3.8 per person

Ph.D. Program

The Department of Biology introduces graduate students to diverse fields of biological science, and provides them with expert guidance to excel in research. The department is invested in training students to become excellent scientists, researchers, science communicators, and instructors. We are a diverse and global community, committed to expanding scientific career opportunities to all. Some of our graduates become academics, whereas others find careers in government, private industry, public policy, or elsewhere (see where recent graduates are now). The ability to communicate ideas and research results clearly and convincingly is key to success in any career.

Experience Biology at Georgetown University

Professors and current students share their perspectives on Georgetown’s cutting-edge biology graduate program.

Potential applicants are urged to identify and contact potential research mentors directly before applying. Please refer to the research page and the list of faculty interested in accepting students . Not all laboratories will have open positions available for a new graduate student in a given year.

Financial Assistance for PhD Students

    students who are accepted into the phd program in biology are guaranteed a minimum of five years of full funding. this funding includes:.

Stipend The stipend for PhD students in Biology is set by the Collective Bargaining Agreement between Georgetown University and the Georgetown Alliance of Graduate Employees ( GAGE ). The current stipend rate is $36,934 for the year. The stipend is paid on a 12-month contract, typically in 26 biweekly paychecks.

Tuition Waiver The Graduate School of Arts & Sciences (GSAS) provides full tuition waivers for all PhD students for the duration of the PhD program.

Health Insurance Health insurance is required. The GSAS provides free student health insurance to all students. The insurance covers doctor’s visits, hospital stays, and prescription drugs. Students may waive this benefit if they prefer a different plan through parents or a spouse.

Dental Insurance The GSAS provides free dental insurance to all students. The insurance covers routine dental care, such as cleanings and exams.

Yates Athletic Facility All students are eligible to use the Yates athletic facility for free. The facility has a gym, swimming pool, and fitness classes.

Parental Leave Graduate student workers are guaranteed six weeks of paid parental leave. This leave can be used to bond with a new child or to care for a sick child.

Medical Leave Graduate student workers are guaranteed six weeks of paid medical leave. This leave can be used to recover from a medical condition or to care for a sick family member.

To learn more about financial assistance for PhD students, please visit the Graduate Funding page. You can read more about what it means to do a PhD in a union-protected graduate program  here

Student Quotes

“Until you start your journey in grad school you might not know what you are missing out on. Other than the course work, grad school has been instrumental in  inculcating important life values and skills such as confidence, time management, survival, and holding personal and professional relationships in higher esteem. ”

“Studying at Georgetown has enabled me to be a lot more confident in my ability to ask questions and chart my own path in scientific research. My advisor and mentors here at Georgetown have been top quality in terms of both academic and personal support”

“Georgetown is a place with an environment conducive to learning and professional growth for prospective scientists working towards their graduate degrees and beyond. In this program, I found mentorship and training that foster the critical thinking required to answer the questions I am interested in. I found the multidisciplinary nature of the graduate student body to be engaging and supportive from the beginning of my journey.”

“The breadth of research that goes on in the Bio department really facilitates a broad understanding of biology from the life of single cells to the life of much larger organisms. It really is something unique that you don’t get at other more specific departments and has really helped me think outside of the box with my own work.”

Graduate Research Opportunities

Learn about the research programs that the department of Biology offers and find out which programs are recruiting graduate students.

Graduate Teaching Opportunities

Find out about the many opportunities that are offered to graduate students to develop teaching and mentoring skills.

Graduate Funding Opportunities

Learn about stipend funding and the additional funds from the GU Graduate School offered to help support graduate dissertation research or travel to meetings.

Doctor of Philosophy (PhD) in Biology

The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of full-time study.

A summary of Biology PhD student expectations by year can be found here . Full details can be found in the Graduate Program Guide .

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program.

How to Apply    Frequently Asked Questions

Learning Outcomes

1. Demonstrate academic mastery in one of three areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

2. Attain research expertise , including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

3. Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

4. Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Coursework Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits.  See the  Graduate School of Arts & Sciences (GRS) Transfer of Credits policy  for more details. 

• 2 semesters of Progress in Research Seminars (2 credits each):

Cell & Molecular Biology:  BI 583 & BI 584 (CM section)

Ecology, Behavior & Evolution: BI 579 & BI 580

Neurobiology:   BI 533 & BI 5834

• 1 semester grant-writing course (2 credits):

Cell & Molecular Biology:  BI 581

Ecology, Behavior & Evolution: BI 671

Neurobiology: BI 581

• 1 semester pedagogy course (1 credit): All Biology PhD students take  BI 697
• 1 quantitative course (3–4 credits): from a list of recommended courses .
• Additional courses vary by specific track below:

Cell & Molecular Biology

1. GRS BI 791/GRS BI 792 Graduate Rotation Credits (2 credits each/4 credits total)

2. GRS BI 753 Advanced Molecular Biology  (4 credits)

3. GRS MB 721 Graduate Biochemistry  (4 credits)

4. GRS BI 735 Advanced Cell Biology  (4 credits)

5. Two electives (8 credits, 500-level and above), see the Graduate Program Guide for recommendations

6. Research credits, CAS BI 925/CAS BI 926  (remaining credits)

Ecology, Behavior & Evolution

1. Six electives  (24 credits, 500-level and above), see the Graduate Program Guide for recommendations

2. Research credits, CAS BI 911/CAS BI 912  (remaining credits)

Ecology, Behavior & Evolution PhD candidate coursework is highly variable. Students, in consultation with advisors, develop a plan of coursework and research. Students are required to take a minimum of 32 credits of coursework. The remainder of the credits should be research.

Neurobiology

2. GRS BI 755 Cellular and Systems Neuroscience  (4 credits)

3. GRS BI 741 Neural Systems: Functional Circuit Analysis  (4 credits)

4. Four electives  (16 credits, 500-level and above), see the Graduate Program Guide for recommendations

5. Research credits, CAS BI 939/CAS BI 940  (remaining credits)

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge . The course provides guidance and training on pedagogy and other aspects of graduate school.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the  Cell & Molecular Biology and Ecology, Behavior & Evolution curricula , this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation & Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the  GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Graduation Timeline

Forms and additional information about PhD graduation can be found on the GRS website .

9 – 12 months before proposed graduation date

• Dissertation Prospectus & approval form due to Graduate Program Specialist for departmental review and submission to GRS

Semester prior to your intended graduation cycle

• Intent to Graduate Form completed online

About 2 months before dissertation defense

• Meet with Graduate Program Specialist, Director of Graduate Studies, and Faculty Advisor to review requirements for defense
• Arrange for Special Service Appointments if committee members are not BU faculty
• Send first draft of dissertation to readers

Once defense date is confirmed with committee

• Reserve room(s) for public seminar and defense

At least three weeks prior to dissertation defense

• Schedule of Final Oral Exam with Abstract Approval due to GRS
• Properly formatted draft of dissertation submitted as PDF to [email protected]

At least two weeks prior to dissertation defense

• Send dissertation to all committee members

At least one week prior to dissertation defense

• Send program information to Graduate Program Specialist

See the Graduate Program Guide for final dates to submit dissertation to ETD

• Submit final dissertation to ETD (online submission)
• Complete electronic signature page and forward confirmation to Graduate Program Specialist along with a copy of your submitted dissertation

MS Degree (En Route to PhD)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam), and has completed 32 credits of graduate-level coursework (not including research) may apply to the Graduate School for an MS degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two: A PhD student who has taken, but has not advanced to candidacy based on the PhD qualifying examination, may still receive an MS degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for a MS degree, and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Interdisciplinary Study Options

Biology PhD students have the option to participate in the Boston University Graduate Program in Urban Biogeoscience and Environmental Health (BU URBAN), the National Science Foundation Research Traineeship Program Understanding the Brain: Neurophotonics (NSF NRT UtB: Neurophotonics), and the Biogeoscience Advanced Graduate Certificate Program . These programs require separate applications in addition to the standard Biology PhD application; those interested in BU URBAN are encouraged pre-apply .

Time Limits

Officially, the PhD must be completed within seven years after the first registration for doctoral study. PhD degrees are conferred in either May, August, or January, as specified on the GRS website . In addition, the PhD candidacy expires after the fifth anniversary of passing the Qualifying Examination. Petitions to extend this deadline are possible at the discretion of the Director of Graduate Studies, the Department Chair, and the Dean of the Graduate School, and can be obtained from the Office of the Graduate School of Arts & Sciences.

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program. PhD students are encouraged to apply for fellowships and grants at funding agencies. All domestic students should apply for NSF Graduate Research Fellowships in the Fall semester of their first or second year. 

Travel Grants may be available to assist students in their travel to professional scientific meetings; students presenting papers or posters on their research will receive first consideration.

Common Types of Funding:

Dean’s Fellowships: These are non-service fellowships allocated to first-year PhD students that do not have immediate teaching requirements.

Teaching Fellowships: These provide a stipend plus full tuition and fees for up to four full courses per semester plus a 2-credit teaching course. Teaching responsibilities usually require approximately 20 hours per week. Full or partial awards may be given.

Doctoral Research Fellowships: These awards are given to students who assist individual faculty with specific areas of research. These Research Fellowships provide a stipend and full tuition. The supervising faculty member determines the specific duties of the Research Fellow.

In addition to the above funding sources, several competitive Department awards and fellowships are available to graduate students in the Department of Biology.

Back to Top

PhD in Biology

The PhD program in Biology is a research-intensive program that also has a strong focus on teaching, designed to produce top scientists and educators with a broad base of knowledge to tackle the most important biological problems of today. This is accomplished through research training, graduate-level courses, seminars, and teaching experience. Entering students are assigned an advisory committee of three faculty members who work with the student to plan a suitable program based on the student's experience and interests.

PhD students in Biology receive full tuition support for six years, a competitive stipend, health coverage, and receive extensive training in pedagogy, outreach, and communication that prepares them for careers in academia, biotechnology, education, and policy. Students are required to serve as teaching assistants for at least two semesters. Most students complete the requirement for teaching experience in the first year by assisting in the introductory undergraduate biology courses for two semesters.

During the first year, students become familiar with research opportunities in the department by doing short research rotations with various faculty members. This experience in expected to lead to the selection of a research focus for the PhD thesis, and it also sets the stage for cross-disciplinary approaches to the thesis topic that is pursued.

First- and second-year students usually take some graduate courses that are selected in consultation with the advisory committee, in addition to one required course: either Biology 243: Topics in Molecular and Cell Biology; Biology 244: Topics in Evolutionary Ecology; or Biology 246: Topics in Physiology and Animal Behavior.

The department faculty are educators as well as researchers, and the graduate students benefit from this expertise. Graduate students serving as teaching assistants have the opportunity to take a course on teaching and pedagogy to improve their skills, or to become involved with upper level courses in their area of expertise. In addition, we offer a one-month summer program for PhD students interested in contributing to course design and honing their teaching skills via GIFT, the Graduate Institute For Teaching .

Research Concentrations

Browse the six concentration areas below for recommended programs of study and relevant courses:

• Biology Education Research
• Ecology, Behavior and Evolution
• Global Change Biology
• Genetics and Molecular Biology
• Developmental Biology
• Physiology, Neurobiology and Animal Behavior

Graduate credit for a course requires a grade of B- (B minus) or better. A list of biology department undergraduate and graduate courses can be found by selecting the Courses button from the top bar.

Throughout the year, graduate students benefit from a variety of seminars on current research that are presented by faculty, fellow graduate students, and invited speakers.

• Skip to Main
• CGSB Faculty
• Affiliated Faculty
• People in History
• Genomics & Systems Biology
• Developmental Genetics
• Molecular & Cellular Biology
• Computational Biology
• Infectious Diseases and the Microbiome
• Evolution, Ecology & Environmental Biology
• Plant Genomics
• Undergraduate Program
• Master's Programs

PhD Program

• Graduate Student Awards
• Course Listings
• Programs & Activities
• Graduate Program Diversity
• Mitra Lecture Series and Fellows
• Conference for Undergraduate Research in Biology (CURB 2024)
• Resource Guides
• Rooms & Calendars

NYU Biology’s PhD program offers training in a broad range of biological research fields, including developmental genetics, genomics and systems biology, molecular and cellular biology, evolutionary biology, and infectious disease. Our dynamic and diverse community of faculty and graduate students engages closely on all aspects of scientific investigation. Strong mentoring relationships throughout the PhD program develop students’ abilities to independently design and perform research, write grant proposals, and communicate their findings effectively to a wide audience. The combination of cutting edge research, world class faculty, and a dedication to mentoring and career building skills for our PhD students defines the PhD program at NYU Biology.

Share this page

Molecular and Cell Biology

Doctoral Program

The Department of Molecular & Cell Biology at the University of California, Berkeley offers a Ph.D. program focused on the molecular mechanisms inherent to life. This program integrates research with a modern training curricula, teaching, and career mentorship. Our Department is highly interdisciplinary - comprising the Divisions of Cell Biology, Development & Physiology, Immunology and Molecular Medicine, Molecular Therapeutics, Biochemistry, Biophysics & Structural Biology, and Genetics, Genomics, and Development – and this is reflected in our students and training. The program is also highly collaborative with related programs and Institutes on campus, thus allowing students the flexibility to explore all aspects of modern biological research. Please click on the links below to learn more about our areas of research on the main department website or use the menu at the top to navigate to areas of interest within the graduate program.

Cell Biology, Development & Physiology

Immunology and molecular medicine, biochemistry, biophysics & structural biology, molecular therapeutics, genetics, genomics, evolution, and development.

Biological Sciences

• Mellon College of Science

Ph.D. in Biological Sciences

The mission of the Ph.D. program in Biological Sciences is to train new generations of scientific leaders who make significant and distinctive contributions to society. The department has established a broad range of resources to enable the students to define and achieve their personal mission. With this broad goal in mind, our specific objectives are to mentor students to:

Become contributing, impactful members of their discipline. This requires:

• Being familiar with literature and history of the field
• Understanding how the discipline has evolved and where it is headed
• Identifying novel and important questions
• Identifying desired personal impact on the field.
• Mastering approaches and techniques
• Appreciating the impact of the discipline on a broader community
• Establishing visible profile in the field

Develop professional skills in :

• Scientific writing (papers, proposals, traditional and social media)
• Oral presentation
• Team management (including interdisciplinary teams)
• Communication, networking, connecting with community of researchers & science professionals
• Time management, life/work balance

Establish career development plans through :

• Establishing sense of scholarly citizenship
• Knowing the types of careers paths following from Ph.D. training
• Developing personal career goals
• Appreciating skills required to achieve career goals and how to align personal training with these goals 

Ph.D. Student Research Videos

Emily simon (linstedt lab), andrew wolff (hinman lab).

• Online Application

" CMU provides an innovative environment, where technology meets deep scientific thinking. I received constant guidance both for my research projects and career path. I had the opportunity to explore different areas and expand my imagination about what is possible to discover in biology, by taking selective courses and consulting faculty with a broad range of expertise. Through presenting at international conferences, I established scientific collaborations, which lead to significant findings and publications, one of which is in the highly regarded journal  Nature . A key perk of being a part of the Department of Biological Sciences is the alumni network of CMU, which consists of overachieving and fearless individuals. As a recent hire of a company started by two CMU alumni, I am excited to continue my pursuit of discovering the unknown and challenging the current limits of experimental sciences ."

" One of the characteristics of CMU that I particularly loved was the extensive levels of collaborations between research groups, even from seemingly unrelated fields. Such exquisite collaborations often further the frontiers of research and being a graduate student at CMU allowed me to lead and be a part of such high quality research. This is primarily due to the excellent faculty at CMU who are always interested in exploring new ideas and are very approachable which gave me immense encouragement  ..... Pittsburgh also provided ample opportunities to pursue my varied hobbies, be it art and music or skiing and rock climbing. This was particularly important for me as it provided a way to switch off from research and rejuvenate. "

" As a prospective student in neuroscience, most of the graduate programs I applied to were neuroscience only programs. One of the biggest reasons I am so glad I chose the Biological Sciences graduate program at CMU, is because I not only gained access to excellent training in the field of neuroscience, but I was surrounded by faculty, post-docs, and other graduate students from many other disciplines. This provided me with different perspectives on my research and enabled me to have access to, learn, and incorporate a variety of techniques from different fields during my graduate experience. It also provided me with experience working as part of an interdisciplinary team, which is a very important part of my job now as I work closely with faculty from mathematics, physics, chemistry, and biology. "

• Graduate Students
• Administration

Skip to content

School of Biological Sciences

• College of Sciences

Search form

Ph.d program in biology.

Our PhD program accepts students possessing Bachelors or Masters degrees who are planning a career in science. The purpose of our PhD program is to prepare highly qualified scientists who have excellent up-to-date training in the fields of biology or bioinformatics, who are actively involved in scientific research, are capable of making significant contributions to their scientific field, possess all the necessary skills for effective oral and written communication with colleagues, and can successfully compete in the job market for postdoctoral and research scientist level positions in academia and science-related industries. We currently have approximately 100 PhD students in the Biology PhD program of the School of Biological Sciences. Of recent PhD graduates, 70% are currently employed as postdoctoral researchers in academia, 9% as government scientists, 9% as industry scientists, 4% as instructors, and 9% are enrolled in further professional training. The average number of journal publications is 3.9 per student and the average number of presentations at conferences around the world is 4.3 per student.

Areas of concentration:

• Ecology, Evolution, and Behavior   including population and evolutionary ecology; community ecology; aquatic chemical ecology; ecological genomics; sensory ecology; evolution of development, behavior, and sociality; biological oceanography; environmental microbiology; theoretical ecology.
• Molecular and Cell Biology   including eukaryotic and prokaryotic cell biology; molecular physiology; molecular biophysics and structural biology; animal, plant, and microbial molecular genetics; human genomics; molecular evolution.
• Computational Biology and Bioinformatics   (with available   M.S.   and interdepartmental   PhD   programs in Bioinformatics) focusing on DNA and protein sequence analysis; comparative genomic analysis; macromolecular structure modeling including protein-protein, protein-nucleic acid, and protein-small molecule interactions; molecular evolution.

Ph.D. Program Overview:

The PhD in Biology is a research-based degree involving deep immersion in a topic within biology with mentoring from an advisor and thesis committee with expertise in the field of study. Coursework is typically completed within the first 2 years (18 credit hours). Students who have previously earned an M.S. degree or taken graduate courses elsewhere can negotiate up to 9 transfer credit hours upon entering our program. Selection of a thesis advisor is made in discussion with our faculty and/or via lab rotations during the first year, and the composition of the thesis committee is established by the student by the end of the first year.

PhD students in the School of Biological Sciences are each supported by a stipend and do not pay tuition, only modest   fees   each semester. Stipend support comes from teaching or research assistantships which complement research training towards the PhD. Whether PhD stipends are earned from research or teaching assistantships after the first year is decided in consultation with a student's thesis advisor and the graduate committee. Our PhD stipend is $33,500. Several fellowships are available which can supplement the PhD stipend. 

Further Funding: Please take some time to review our   funding opportunities and fellowships .

Choosing between the Masters and Ph.D. Program: Our graduate program is primarily focused on the PhD which prepares students for careers in scientific research and employment in academia, industry, or government. We encourage potential applicants to the PhD program to directly contact individual   faculty   members whose research may appeal to them to discuss research interests and future opportunities. Students who are not yet sure of their interest in scientific research or are interested in other kinds of professional development should consider the M.S. degree in   Biology   or Bioinformatics . However, we accept only a small number of M.S. Biology students each year - typically those in special career circumstances with a prior relationship to our program and faculty. Admission decisions are made by our graduate committee (composed of Biology faculty) in consultation with all of the faculty in the School of Biological Sciences.

Georgia Tech Resources

• Offices & Departments
• News Center
• Campus Calendar
• Special Events
• Institute Communications
• Visitor Resources
• Campus Visits
• Directions to Campus
• Visitor Parking Information
• GTvisitor Wireless Network Information
• Georgia Tech Global Learning Center
• Georgia Tech Hotel & Conference Center
• Barnes & Noble at Georgia Tech
• Ferst Center for the Arts
• Robert C. Williams Paper Museum

College of Sciences Social Links

Biology related research centers.

• Aquatic Chemical Ecology Center
• Center for Biologically Inspired Design
• Center for Integrative Genomics
• Center for NanoMAD
• Center for Ribosomal Evolution and Adaptation
• Center for the Study of Systems Biology
• Integrated Cancer Research Center

Georgia Institute of Technology School of Biological Sciences 950 Atlantic Drive Atlanta, GA 30332 Office: 404-894-3700 Fax: 404-894-0519

Georgia Institute of Technology North Avenue, Atlanta, GA 30332 404.894.2000

• Emergency Information
• Enable Accessibility
• Legal & Privacy Information
• Human Trafficking Notice
• Title IX/Sexual Misconduct
• Hazing Public Disclosures
• Accessibility
• Accountability
• Accreditation

© Georgia Institute of Technology

• Skip to Content
• Catalog Home
• Institution Home
• Graduate Catalog /
• School of Arts & Sciences /

Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2024 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

Sample Plan of Study

Print Options

Print this page.

The PDF will include all information unique to this page.

A PDF of the entire 2024-25 catalog.

A PDF of the 2024-25 Undergraduate catalog.

A PDF of the 2024-25 Graduate catalog.

• Department of Biology and Biochemistry

Graduate Programs

• Prospective Students

Ph.D. Programs

The Department of Biology & Biochemistry offers Ph.D. degrees in Biochemistry and in Biology. The Ph.D. program in Biology has two degree tracks: the Cell and Molecular Biology degree track, and the Ecology and Evolution degree track.

Biochemistry

Faculty and graduate student research focuses on biochemical processes at the subcellular and macro-molecular levels and encompasses a variety of fields and methodologies. Areas of study include macromolecular structure and function as elucidated by nuclear magnetic resonance; X-ray crystallographic and spectroscopic techniques; enzyme reaction mechanisms; genomics; computational methods in molecular biology; computational biochemistry/biophysics; computer-aided drug design; signal transduction; neurochemistry; ion channel structure and function; the role of RNA in molecular evolution; the structure and function of virulence factors; and biotechnology.

• View requirements

Cell and Molecular Biology

The faculty and students in this program share common interests in understanding the molecular mechanisms which drive both fundamental cellular processes and the developmental processes of morphogenesis, cell differentiation and gene regulation. The strength of the program is the diversity of the biological systems under study, which stimulates extensive exchange and collaboration between the various groups. Faculty expertise spans the disciplines of cell and developmental biology, molecular biology, physiology, microbiology, neuroscience, immunology, and genetics.

Ecology and Evolution

This program blends knowledge and methodology from diverse biological disciplines to better understand ecological and evolutionary processes operating at multiple scales—from molecules to individuals to societies to communities. Current research programs include experimental evolution, evolution of development, evolutionary genetics, behavioral ecology, community ecology and evolutionary bioinformatics in systems ranging from bacteria to ants, from protists to grasses. Faculty conduct studies in natural habitats including the Colorado plateau, and coastal salt marshes, and in artificial systems such as petri dishes and theoretical models.

Financial Assistance

The Department of Biology & Biochemistry believes that high-quality graduate studies require a commitment to high-quality research. As a result, our graduate students receive financial support sufficient to provide a modest standard of living that enables them to make a full-time commitment to their graduate training. Some of the types of financial support available to students are listed below. Additional financial assistance may also be available from the College of Natural Sciences and Mathematics and the University of Houston Graduate School .

Teaching Assistantships (TAs)

TAs are the main source of support for first-year students but are available in subsequent years for students not supported by grant funds. TAs will be provided a salary of $2,194.66/month (U.S. $26,335.92 per year). This level of support is sufficient for international students to obtain an F1 visa.

Research Assistantships (RAs)

RAs are the main source of support for students after their first year in the program. RA support is provided through grants to the lab the student joins to conduct graduate research. RA support is currently $2,194.66/month (U.S. $26,335.92 per year).

Graduate Tuition Fellowship (GTF)

Students supported as TAs or RAs are eligible for tuition fellowships to cover their mandatory tuition and fees. These fellowships provide the students with approximately $20,000/year to pay for mandatory tuition and fees. See more information on eligibility criteria .

Non-resident Tuition Waiver

Out-of-state students and international students employed as TAs or RAs receive a waiver of the additional tuition charged to non-residents.

Presidential Fellowship

All applications submitted for admission to the Biology & Biochemistry Graduate Program are reviewed by our Graduate Recruiting and Admissions Committee. This committee is comprised of a group of faculty from each division of the department. Once admitted to the program, accepted applicants are further evaluated for the Presidential Fellowship. The criteria for evaluation are as follows:

• Appropriate academic coursework in the major
• Outstanding grades in previous university coursework
• Demonstrated research ability or potential for research excellence.

If awarded, the student receives $2,000/per year for the first two years. This fellowship is in addition to the monthly stipend and Graduate Tuition Fellowship given to all students admitted to our graduate program. Students must meet minimum full-time enrollment (9 hours) and a cumulative 3.00 GPA to maintain the fellowship each semester it is held.

Houston Livestock Show and Rodeo Scholarship

This scholarship is awarded to outstanding students enrolled in our graduate program. The maximum award is $2,000/year. Recipients must be Texas residents and citizens or permanent residents of the United States. Students apply annually on the UH Foundation website.

Medical Insurance

In addition to their stipend, graduate students employed as TAs or RAs receive $150/month for health insurance. For more information about the student health insurance plan endorsed by and designed especially for the University of Houston, please see Student Health Insurance .

Cost of Living

Houston has a relatively low cost of living compared to most major U.S. cities and many low-cost apartments and houses are available.

Admission Requirements

The minimum entrance criteria for doctoral graduate studies in the Department of Biology & Biochemistry are as follows:

• Completion of a baccalaureate degree (B.S.) with a major in Biology, Biochemistry, or an equivalent discipline. You can apply to our programs before you complete your degree, provided you graduate before you enter the program. ( NOTE: A prior M.S. is not a requirement to apply to our Ph.D. programs.)
• Grade Point Average ≥ 3.0 / 4.0 (overall or for the last 60 hours of coursework completed).
• The GRE is no longer required by our Ph.D. programs . If you decide to submit GRE scores, UH's Institutional Code is 6870.
• Informative, coherent and well-written statement of purpose .
• Three strong letters of recommendation .
• English Language Proficiency Requirement. All applicants, regardless of citizenship status, must demonstrate proficiency in English to obtain admission. To fulfill this requirement, applicants must satisfy one of the following criteria: a) Bachelor's degree (or higher) earned from a regionally accredited U.S. institution or at an institution at which English is the medium of instruction in the following countries: Australia, New Zealand, South Africa, the Bahamas, the United Kingdom, Ireland, Jamaica, Liberia, Trinidad, the Virgin Islands, Antigua and Barbuda, Dominica, Saint Lucia, Saint Vincent and the Grenadines, Barbados, Grenada, Turks and Caicos, and English-speaking Canadian provinces. b) TOEFL. The minimum TOEFL score required is 79 for the internet-based test. The minimum TOEFL score for the new revised paper-based exam is: Reading 20, Listening 20, and Writing 20. TOEFL scores must be received directly from Educational Testing Service (ETS). UH's Institutional Code is 6870. c) IELTS. The minimum IELTS score required is an overall score of 6.5. The testing agency should mail the official results directly to UH. No electronic IELTS are accepted.  d) Duolingo. A minimum score of 105 is required.

Use the online application to submit all your documents electronically. Your references will be automatically contacted to submit their letters of recommendation. Please follow the instructions on the UH Graduate School Application page.

Tips for Applying

What we look for.

We seek to admit students who show a strong record of academic achievement and a high level of motivation and interest. Your record of academic achievement and ability is conveyed by your transcripts, GPA, and Graduate Record Exam (GRE) scores, as well as your letters of recommendation. Your level of motivation and interest is conveyed by your personal statement and letters of recommendation. We will evaluate your application on the basis of your transcripts, test scores (GRE scores for everyone, TOEFL/IELTS scores for foreign students), your personal statement, and the letters of recommendation.

Transcripts and GPA

Successful applicants to our program generally have GPA's of 3.00 or higher. However, a student with a high GPA and a transcript with lots of non-rigorous courses may not be viewed as favorably as a student with a somewhat lower GPA who has taken courses that are essential preparation for graduate work (such as Genetics, Cell Biology, Biochemistry, Evolutionary Biology, etc.). It is not essential to have all of these" foundation" courses before you start graduate school, but if you do not have most of them, you will not be well prepared for graduate school.

The GRE provides information regarding your overall academic ability. You are more likely to do well on the GRE if you prepare for the exam. Preparation guides and practice tests are available at most college bookstores.

Your school may provide assistance in preparing for the GRE; check with your career or academic counseling office.

TOEFL/IELTS

This exam is required for all applicants who have not obtained a prior university degree from an institution where English is the medium of instruction (see list of exempt countries and English Language Proficiency Requirements ). These scores must be officially reported to the University before we can admit you to the program or offer financial support.

Statement of Purpose

Your personal statement is your opportunity to tell us why you want to join our graduate program and what your long-term goals and interests are. You do not have to know exactly what you want to do, or what scientific questions you want to pursue, but you should tell us what excites your interest or curiosity. Be specific. Your statement is also a chance to discuss any aspect of your application (such as academic history) that you feel warrants further explanation.

If there is a reason for your low GPA (a bad semester due to personal difficulties, for instance), you can provide a brief explanation in your personal statement. High GRE scores can make up for a low GPA, and a high GPA can balance out low GRE scores. In some cases, research experience and strong letters of recommendation can make up for low grades and low GRE scores.

Letters of Recommendation

You will need 3–4 letters of recommendation. Most or all should be from your professors, and the letters should come from people who know you personally as well as your academic performance. Make sure your letter writers know your academic record, reasons for wanting to go to graduate school, and long-term goals.

Contacting Faculty

You can contact individual faculty members in our department whose research is of interest to you, either before or after sending your application. Faculty interests and contact information are available on this Faculty Profiles webpage.

Find more information about the application process on the How to Apply page on the NSM website.

• Open for Applications: June 1
• Deadline: September 1  
• Open for Applications: August 1
• Deadline: January 1  

For More Information

Contact: Rosezelia Jackson 713-743-2633 [email protected]

Secondary Menu

Doctoral degree requirements, general philosophy of the ph.d.  program.

Biology is the most diverse of all the disciplines in the natural sciences. Consequently, the allied fields to which the various subdisciplines in Biology share natural affinities differ. For example, molecular biology makes connections with physical chemistry, biomechanics with engineering and physics, and ecology and evolutionary biology with statistics, mathematics, geology, and atmospheric sciences. The goal of the Biology Doctoral Program is to train young scientists who:

• excel at research and teaching in their own subdisciplines;
• demonstrate competence in fields allied to their subdisciplines; and
• display a breadth of knowledge in Biology as a whole. 

Trajectory through the program

• At the beginning of your Ph.D.: you will be assigned a temporary advisor (typically the sponsor of your admission).
• Before registering for the first semester: discuss with your advisor which courses you should take. As a first-year student, you are encouraged to interact with other faculty in your research area to ask them about courses you should take before the preliminary exam—these faculty may very well become members of your Ph.D. committee.

The philosophy of the department is that you need not spend a great deal of time in coursework.  You should take only the courses that fill gaps in areas that will be needed in your research, and spend most of your time starting your research.

• During the first three semesters: take up to three tutorials with different faculty members in the department. The tutorial requirement is waived as soon as you declare an advisor. Tutorials may involve laboratory work, directed reading in the primary literature, greenhouse or field studies, mathematical or computer modeling, or any other activity that would assist you in identifying a suitable dissertation topic. Other goals of the tutorials are to expose students to the diversity of faculty research interests in their specialty and to help them to identify an appropriate advisor and dissertation committee. 
• By the end of the 3 rd semester, you must choose an advisor and the advisor must convene an initial meeting of the dissertation committee. Dissertation committees will consist of at least four faculty members, one of which will represent your minor (see below). The goals of the initial meeting are to assess the novelty and feasibility of your proposed dissertation topic, to ascertain whether you need to take any additional courses to demonstrate preparedness for the preliminary exam, and to decide the format of the written dissertation proposal (see below).  If you fail to meet with your committee before the end of the 3rd semester, the Director of Graduate Studies (DGS) will notify you in writing, and will discuss with you and your advisor ways to resolve any difficulties. 
• Preferably by the end of the 4 th semester, but certainly by the end of the 6 th semester, you must pass an oral preliminary examination to establish candidacy for the Ph.D. One week prior to the exam, you must present a written essay describing your proposed dissertation research to your committee. The exam itself will cover the specific research areas addressed in your proposal, but will also test your depth of knowledge in your area of specialty and your breadth of knowledge in Biology as a whole.
• When the dissertation research is completed, you will present the written dissertation to your committee two weeks prior to the dissertation defense (the dissertation also has to be submitted to the Grad School two weeks before the defense). You are also required to present your results in a seminar. You are expected to complete the degree requirements as soon as possible, but students making progress toward their degree will be considered to be in good standing through the 12 th semester. 

You are required to declare a minor at the time of the initial meeting with your dissertation committee. At least one member of the dissertation committee must represent the minor field, and will be charged with assessing your knowledge in the minor during the preliminary exam.  The goal of the minor is to ensure that you acquire a breadth of knowledge beyond your immediate specialty, either in a different area of Biology or in an allied field. Dissertation committees have the freedom to determine the specific nature of the minor, to tailor it to your needs, but minors are subject to approval by the Director of Graduate Studies and the Graduate Affairs Committee.

The minor may:

• lie entirely outside Biology (e.g., Statistics, Mathematics, Computer Science, Chemistry, Engineering, Atmospheric Sciences, Geology);
• require the student to acquire knowledge about a group of organisms that differs from those on which the student’s dissertation research focuses; or
• represent a biological subdiscipline that is distinct from the student’s own subdiscipline.  For example, a student whose advisor is in the Evolution, Ecology, and Organismal Biology sub-department might choose Developmental, Cellular, and Molecular Biology (DCMB) as a minor, with appropriate representation by a DCMB faculty member on the preliminary exam and dissertation committees. A student in ecology might choose evolutionary biology as a minor.

Scholarly productivity

To become recognized as independent scientists and to have an impact on their fields, graduate students must communicate their findings to their colleagues. Therefore, all students are strongly encouraged to begin submitting the results of their research to refereed journals as soon as possible (ideally well before the dissertation is completed). In order to give their committee members an opportunity to comment on them, manuscripts that are intended to be included in the dissertation will be given to all committee members at least two weeks prior to submission. You should submit copies of papers accepted for publication to the Director of Graduate Studies, to be included your file. 

You will be required to serve as a teaching assistant for two semesters. Teaching is not simply a mechanism to provide financial support to graduate students. Rather, it is an integral part of your professional development as both an educator and a researcher.

Annual evaluation of student progress

• After your 1 st year you will meet with your committee annually.
• In your 2 nd year, your first committee meeting serves as the annual committee meeting.
• In your 3 rd year, your Preliminary Exam will serve this purpose if you take it in the Fall.

You MUST have an annual progress meeting with your committee each Fall, UNLESS:

• you are in your 1 st or 2 nd year;
• you are in your 3 rd year and are taking your prelim this Fall (not Spring, and not in your 2 nd year); or
• you are defending your dissertation this year.
• Prior to November 15, you are expected to present to your committee a written report on the progress made over the previous year, and on any difficulties encountered, as well as a plan for completing the dissertation.
• Before the end of the final exam period in December, your advisor will then convene a meeting of the committee to discuss the progress report with you.
• By January 1, your advisor must send a letter to the Director of Graduate Studies summarizing this discussion, providing the committee’s evaluation of your progress, and recommending whether you should receive continued financial support from the department. The progress report you submitted to your committee should be attached to this letter. All faculty members not on your committee will also be able to submit to the Director of Graduate Studies any written commentary on the student’s performance (e.g., as a student or teaching assistant in a course taught by that faculty member).  
• After January 1, the Director of Graduate Studies, in consultation with the Graduate Affairs Committee as needed, will review the written materials to determine whether you shall be granted departmental support for the following academic year. You will receive a letter from the DGS if any remedial measures need to be taken. Your written progress report, your advisor’s letter to the DGS, any letters from faculty not on your committee, and any letters from the DGS will be placed in your folder as a record of your annual progress and evaluation. 

Committee Meetings, Exams, and Defenses during Summer

Scheduling committee meetings, preliminary exams, and dissertation defenses during the summer is strongly discouraged, because faculty members typically have research or other travel plans that must take precedence during this time. Summer meetings, exams, and defenses can only be scheduled by agreement of all members of the committee. In accordance with Graduate School regulations, such meetings can only be held while the summer semester is actually in session, and the student must be registered for the summer semester.

Note: A downloadable version of the information above is available:

• Duke Biology’s Mission Statement
• AJED Annual and Semester Reports
• AJED Meeting Notes
• Biology Cultural Association (BCA)
• Inclusion, Diversity, Equity, and Antiracism Committee (IDEA)
• Learning from Baboons: Dr. Susan Alberts
• Extremophiles and Systems Biology: Dr. Amy Schmid
• How Cells Manage Stress: Dr. Gustavo Silva
• Predator-Prey Interactions in a Changing World: Dr. Jean Philippe Gibert
• Exploring the Extracellular Matrix: Dr. David Sherwood
• Cell Division's Missing Link: Dr. Masayuki Onishi
• Listening in to Birdsong: Dr. Steve Nowicki
• Biogeochemistry as Ecosystem Accounting: Dr. Emily Bernhardt
• Building a Dynamic Nervous System: Dr. Pelin Volkan
• Investigating a Key Plant Hormone: Dr. Lucia Strader
• Imagining Visual Ecology: Dr. Sönke Johnsen
• Outreach Opportunities Across the Triangle
• Job Opportunities
• Location & Contact
• Frequently Asked Questions
• Learning Outcomes
• Major Requirements
• Anatomy, Physiology & Biomechanics
• Animal Behavior
• Biochemistry
• Cell & Molecular Biology
• Evolutionary Biology
• Marine Biology
• Neurobiology
• Pharmacology
• Plant Biology
• Minor Requirements
• Biology IDM
• List of Biology Advisors
• Guide for First-Year Students
• Transfer Credit
• Application & Deadlines
• Supervisor & Faculty Reader
• Thesis Guidelines
• Honors Poster
• Past Student Projects
• Study Away Opportunities
• Finding a Research Mentor
• Project Guidelines
• Getting Registered
• Writing Intensive Study
• Independent Study Abroad
• Summer Opportunities
• Departmental Awards
• Biology Majors Union
• Commencement 2024
• Trinity Ambassadors
• Degree Programs
• Ph.D. Requirements
• How to Apply
• Financial Aid
• Living in Durham
• Where Our Students Go
• Milestones Toward Ph.D.
• Graduate School Fellowships
• Useful Resources
• Concurrent Biology Master of Science
• En Route Biology Masters of Science
• Form Library
• Mentorship Expectations
• On Campus Resources
• Fellowships & Jobs
• Meet Our Postdocs
• Department Research Areas
• Research Facilities
• Duke Postdoctoral Association
• All Courses
• Biological Structure & Function Courses
• Ecology Courses
• Organismal Diversity Courses
• Alternate Elective Courses
• Primary Faculty
• Secondary Faculty
• Graduate Faculty
• Emeritus Faculty
• Graduate Students
• Department Staff
• Faculty Research Labs
• Developmental Biology
• Ecology & Population Biology
• Neuroscience
• Organismal Biology & Behavior
• Systematics
• Research Articles & Papers
• Botany Plot
• Field Station
• Pest Management Protocols
• Research Greenhouses
• Centers/Research Groups
• Biology Writes
• Alumni Profiles
• For Current Students
• Assisting Duke Students

Department of Molecular Biology

Graduate Program

The graduate program in the Department of Molecular Biology fosters the intellectual development of modern biologists. We welcome students from a variety of educational backgrounds, and offer an educational program that goes well beyond traditional biology.

The molecular biology department at Princeton is a tightly knit, cohesive group of scientists that includes undergraduate and graduate students, postdoctoral fellows and faculty with diverse but overlapping interests. Graduate students have a wide choice of advisers, with a broad spectrum of interdisciplinary interests and research objectives.

The graduate program offers each entering student the opportunity, with the help of faculty advisers, to design the intellectual program that best meets his or her unique scientific interests. Each student chooses a series of research rotations with faculty members in molecular biology and associated departments (chemistry, computer science, ecology and evolutionary biology, engineering, physics and psychology). Entering students, with the aid of the graduate committee, select core and elective courses from a large number of offerings in a variety of departments and disciplines. This combination of a cohesive department, one-on-one advising, and individualized programs of course work and research provides an ideal environment for graduate students to flourish as independent scientists.

Learn more about the department of Molecular Biology admission requirements, application process and financial support.

Explore the requirements for completing the Molecular Biology graduate program.

Molecular Biology graduate courses and cross-listed courses offered for the most recent academic year.

Our Molecular Biology graduate program offers a diverse range of concentrated areas to advance knowledge and address complex biological challenges.

The graduate M.D./Ph.D is done in partnership with the Robert Wood Johnson Medical School and the Rutgers University (New Brunswick) Graduate School of Biomedical Sciences.

The training grant has fostered an interdisciplinary life sciences research community at Princeton by supporting second and third-year students in the Department of Molecular Biology and genetics research labs.

Information for Graduate Students

Our Molecular Biology graduate program offers a dynamic and supportive environment for all students.

• Life in the Department
• Graduate Student Committees
• Career Resources
• Student Life
• Incoming Graduate Students

Graduate News

Research Areas

Diverse in research interests

Navigation mit Access Keys

• Direkt zur Startseite
• Direkt zur Navigation
• Direkt zum Inhalt
• Direkt zur Kontaktseite
• Direkt zur Sitemap
• Direkt zur Suche

International PhD Program

PhD Program

Biozentrum International PhD Program

Phd students can expect an extraordinary wealth of research expertise, a wide range of courses, intensive individual mentoring, an international community of researchers from over 40 countries and a lively student life. .

Research at the Biozentrum embraces a wide range of research fields . Yet the  31 research groups  also share a main focus: the strong interest to understand how molecules and cells create life. We have studied this mystery for more than 50 years and are one of the leading Life Sciences institutes in the world, spawning several Nobel laureates.

As a PhD student at the Biozentrum, you will become part of an international community. More than 130 young scientists from around the world are currently doing their PhD here. You will engage in an independent research project under the supervision of a Biozentrum faculty member and a PhD Advisory Committee. Apart from a strong mentoring program, you will also benefit from the open and collegial atmosphere that encourages creativity and critical thinking. 

To find out first hand from a student how it is to do a PhD at the Biozentrum, read the interview with Talia Ulmer.

"During my master’s in Neurobiology a lot of really exciting questions came up and as research is really fun for me, I decided to do a PhD and continue my research for a few more years”, explains PhD student Talia Ulmer.  > to the interview

Graduate Teaching Program Besides the practical training in the latest methods and techniques in modern-day molecular biology research, you will profit from the rich, interdisciplinary  Graduate Teaching Program  offered in collaboration with scientists from the Friedrich Miescher Institute (FMI), the Department of Biosystems Science and Engineering (D-BSSE) of the ETH Zurich, the Department of Biomedicine (DBM), and the Swiss Tropical and Public Health Institute (SwissTPH). 

The program includes lecture cycles in Infection Biology, Neuroscience, Growth and Development, Structure and Function of Macromolecules, Computational and Systems Biology, Molecular Biology, Molecular Medicine and also training in practical skills such as "How to be a scientis", scientific communication, imaging, proteomics, genomics and biophysical techniques. 

Community and career development activities As a PhD student, you will also regularly attend scientific meetings and organize, together with your colleagues, your own seminar series,  PhD Club , career guidance events and an annual two-day  PhD Retreat  . 

To promote scientific exchange and interdisciplinary teamwork between the research groups, we organize the weekly Discovery Seminars , which are followed by a get-together, as well as a symposium or a multi-day retreat every year. These offer young scientists, in particular, a platform to gain experience in presenting their research findings. 

Application We are looking for talented and motivated graduates, with a background in biological or biomedical sciences, physics, chemistry, mathematics or computer science. You can either apply

• directly to one of our 31 research groups or
• for a  Biozentrum PhD Fellowship

Information on admission, duration, etc., a checklist and other documents can be found under key facts here . If you have any questions, please contact the PhD Student Office

Geschäftsordnung Biozentrum International PhD Program

This website uses cookies to collect data from users and improve functionality. By continuing to browse the site you are agreeing to this. Receive more information by reading our privacy policy.

The PhD program in Biology provides both a broad background knowledge base and an in-depth study of a specialized area of biology. Students can choose to pursue a concentration in Cell and Molecular Biology or Molecular Microbiology.

In The News

Flashes of Insight

The PhD program in Biology emphasizes close interaction between graduate students and faculty in developing the intellectual and experimental skills required for creative independent research. To be considered for admission to the doctoral program, a student must have a bachelor’s or master’s degree, preferably in a biological science. Individuals with degrees in other natural sciences may be considered, depending on their interests and background.

A minimum GPA of 3.000 for all graduate work is required for the award of the PhD degree. All regulations of the Graduate School regarding maintenance of academic standing apply.

The Cell, Developmental, and Regenerative Biology Group studies questions related to organismal development, aging, regeneration and repair. Our unifying theme is an interest in discovering how organisms build and rebuild tissues during development, after injury, and during aging. Studies in worm, fish, axolotl, mouse, and human tissue culture are providing insights into the development, function, and aging of multiple organ systems. We use genomic and proteomic analyses, and biological imaging approaches, to understand the roles and regulatory networks underlying tissue development and repair.

Biomechanics, Neurobiology, and Behavior Group pursues distinct interests, with a special emphasis on motor learning and control, biomechanics, circadian rhythms, neurogenesis and neural regeneration. Animal models run the gamut from jellyfish and termites to fish and human subjects. Research threads include the social behavior of insects, the neural architectures underlying rhythms, movement and goal-directed behaviors in humans and model organisms, and adult neurogenesis and neuronal regeneration in teleost fish.

Molecular Microbiology Group is interested in understanding the rise of multidrug resistant pathogens. Hence the focus of this group intersects strongly with that of the University in the areas of health and security. The Molecular Microbiology group investigates the bases of dormant persister cells, uncultured bacteria, composition and function of the human and environmental microbiomes, mechanisms underlying mutagenesis, regulation of the DNA damage response, and bacterial development including biofilm formation. Further strengths include the development of platforms for the discovery of new antimicrobials.

• Concentrations in Cell and Molecular Biology or Molecular Microbiology
• Bachelor’s and Advanced-degree entry are possible
• Boston residence requirement of one year of full-time graduate work on campus

Our graduates pursue careers within academia and beyond.

• Booz Allen Hamilton
• Albert Einstein College of Medicine
• Agios Pharmaceuticals
• Masschusetts General Hospital

Application Materials

Application.

• Application fee – US $100
• Unofficial transcripts for all institutions attended (Official transcripts required upon acceptance of admission offer)
• Personal statement
• Three letters of recommendation
• GRE General test – Not required
• Proof of English Proficiency for all applicants

Our priority deadline is December 1 with rolling admissions until March 15, based on space available.

We only admit degree program students for fall semesters.

• Program Website

Request Information for PhD in Biology

Jump to navigation

University of Washington Links

• College of Arts & Sciences
• Directories

• Graduate Students
• Fields of Interest
• Publications
• Opportunities
• Undergraduate
• Postdoctoral Fellows
• Life Sciences Complex
• Recent News
• Newsletters

You are here

Phd program, interested in applying this cycle, rsvp for our information session in november .

Graduate students in the Department of Biology at the University of Washington receive a commitment from the entire department to support their education and research.  Our students make excellent use of the many opportunities available to them, as evidenced by the extraordinary number of fellowships and awards received, the successful publication of high impact papers, and their regular contributions at national and international meetings.  Our students receive additional training by participating in departmental governance, curriculum development, faculty recruitment and other activities. Together these factors combine to promote an unusual degree of professionalism and community spirit.

Member of the Team

By joining the Biology Department at the University of Washington, you will become a part of a world-class research institution.  Here, graduate students are treated as colleagues by a committed faculty who work to mentor students, to provide opportunities to develop research interests, to network both on and off campus, and to publish and present their own research.

You will find a collaborative environment at the University of Washington, with faculty engaging in research with the lab next door, researchers across campus, and teams halfway across the world.  Our graduate students are encouraged to collaborate similarly, seeking additional mentors and colleagues throughout the world who compliment their research interests.

Timeline to Completion

• Grad Spotlight
• Undergraduate Opportunities
• Advising & Appointments
• Awards & Scholarships
• Career & Postgrad
• Departmental Honors
• Handouts & Forms
• Internships
• Registration & classes
• Research Opportunities
• Transfers & Postbacs
• Awards & Fellowships
• Ph.D Program
• Prospective Postdocs
• Postdoc Resources
• Course Catalog
• Docent program
• Volunteering
• Backup Freezers
• Information
• Biology Study Area
• Tri-Beta Assistance
• Burke Museum
• Friday Harbor Labs
• COVID-19 information
• Graduation Reception
• Medicinal Herb Garden
• Nature's Depths
• Support UW Biology
• Dept Culture Office Hour
• Diversity and Equity Committee
• Join UW Biology
• Departmental committees
• Mentoring resources
• Organization chart
• Virtual tour

School of Dentistry

• About Our School
• Points of Pride
• Our Mission & History
• Department and Divisions Overview
• Department of Developmental and Surgical Sciences
• Department of Diagnostic and Biological Sciences
• Department of Primary Dental Care
• Department of Restorative Sciences
• Faculty Emeriti & Faculty Ad Honorem
• Diversity, Equity and Inclusion Overview
• Building a Diverse, Equitable and Inclusive Community
• Diversity, Equity and Inclusion Resource Library
• Faculty Development
• Degrees & Programs Overview
• Doctor of Dental Surgery
• Dental Therapy
• Dental Hygiene
• Graduate, Specialty and Advanced Education Programs Overview
• Advanced Education Scholarships
• Endodontics
• Graduate Program in Oral Biology
• Master of Science in Dental Hygiene
• Master of Science in Dentistry
• Oral & Maxillofacial Surgery Training Program
• Orofacial Pain
• Orthodontics
• Pediatric Dentistry
• Periodontology
• Prosthodontics
• Oral Health Educator Certificate Program
• Dental School Preparation Programs Overview
• Discover Dental School
• Open House: Explore Dental Careers
• Community Outreach Experience
• Degree & Education Verification
• Scholarships & Student Support
• Continuing Dental Education Upcoming Courses
• Find a Course
• Dentistry Magazine
• White Coat Ceremony 2024
• Research & Discovery
• Overview of Student Research and Training Opportunities
• Summer Research Fellowship Program
• Minnesota Craniofacial & Oral Health Research Experience
• Faculty Labs
• Oral Health Clinical Research Center Overview
• Faculty and Personnel
• Current Studies
• Data Access and Informatics Consulting
• How to contact a biostatistician from Biostatistical Design and Analysis Center (BDAC)
• Clinical and Translational Science Institute (CTSI): Resources, Services and Funding
• Research and Innovation Office (RIO)
• Minnesota Dental Research Center for Biomaterials and Biomechanics
• Orofacial MRI Center
• Alumni & Donors Overview
• Alumni Adjunct Faculty
• Learners to Leaders
• Connect with Students
• School of Dentistry Alumni Society
• Giving Opportunities Overview
• Ways to Give
• Estate Giving
• Dean's Circle
• Donor Recognition
• Give to the Max Day
• Memorial Scholarships
• Distinguished Alumni Awards
• Dental Hygiene Alumni
• Alumni Weekend
• Campaign priorities
• School of Dentistry Alumni Society Overview

PhD in Oral Biology Program

The PhD in Oral Biology emphasizes basic science research and training to develop independent investigators.

The four-year PhD in Oral Biology encourages students to focus in one of five areas: biomaterials and biomechanics, epithelial biology and carcinogenesis, microbiology and immunology, sensory neuroscience or bone biology, craniofacial development and tissue engineering. 

The program is designed to ensure that graduates will develop the capability to initiate independent research programs in important areas of oral biology. Most students execute projects that advance mechanistic understanding.

Program at a Glance

Joining the program.

We welcome a diverse pool of applicants, including international students. 

Applicants must possess a bachelor's degree or equivalent, or a DDS/DMD degree or equivalent (international dental degrees (BDS, etc.) are acceptable).

PhD applications are due December 1 for the program beginning the following fall. Applicants are not admitted during the spring semester.

Applications must be completed online and should include a personal statement, a research essay, a diversity statement, CV or resume, proof of English language proficiency, letters of recommendation and transcripts. 

Application Instructions

+ Program Requirements

+ Curriculum

The first year of the PhD program consists primarily of coursework. Students select courses with their advisor's approval from a core curriculum recommended by the Graduate Faculty for each area and their minor program. The core curriculum provides students with a working knowledge of the major concepts and research paradigms in that scientific area, a working vocabulary and the basis for continued learning. 

In the second year, students complete all coursework and the written and oral preliminary exams. The written and oral exams capture the student’s ability to think critically about the field and the application of logical experimental designs to test hypotheses and answer questions. Upon completion of this two-part preliminary examination of the research proposal, the student will work largely on the dissertation research project through month 45 in residence.

Months 45 through 48 will be used for dissertation writing. Students must also present a public seminar describing their dissertation research (which is attended by the final oral exam committee) no later than six months before defense of the thesis. The dissertation will be defended in another public seminar in month 48.

• DDS/PhD Program
• PhD Program
• Current Students
• Oral Biology Faculty
• Faculty and Student Resources
• About the Program
• Orthodontics Overview
• Pediatric Dentistry Overview
• Periodontology Overview
• Prosthodontics Overview
• Research & Mentorship

Computational Biology PhD

The main objective of the Computational Biology PhD is to train the next generation of scientists who are both passionate about exploring the interface of computation and biology, and committed to functioning at a high level in both computational and biological fields.

The program emphasizes multidisciplinary competency, interdisciplinary collaboration, and transdisciplinary research, and offers an integrated and customizable curriculum that consists of two semesters of didactic course work tailored to each student’s background and interests, research rotations with faculty mentors spanning computational biology’s core disciplines, and dissertation research jointly supervised by computational and biological faculty mentors.

The  Computational Biology Graduate Group  facilitates student immersion into UC Berkeley’s vibrant computational biology research community. Currently, the Group includes over 46 faculty from across 14 departments of the College of Letters and Science, the College of Engineering, the College of Natural Resources, and the School of Public Health. Many of these faculty are available as potential dissertation research advisors for Computational Biology PhD students, with more available for participation on doctoral committees.

• Cornell University Home
• College of Agriculture & Life Sciences Home

Molecular Biology and Genetics

Prospective Students

The first year in BMCB, GGD, and Biophysics includes an orientation week, rotations in labs with the goal of selecting a lab, and first year assessments. 

Incoming students are required to attend Department of Molecular Biology & Genetics (MBG) Orientation, which generally takes place the week before classes start.  During this time, students will complete required training and paperwork necessary for their program.  It is our goal to have administrative requirements completed before the start of classes in order to allow students to concentrate on the academic part of their program.

Included in the mix with administrative requirements and training are activities designed to welcome you into the MBG and the respective graduate fields.  The Graduate Student Associations will have social programs for incoming students.  The MBG Annual Picnic is held at the beginning of the fall semester and is a good way to meet the students, staff, and faculty in an informal, non-lab setting.

All graduate students in MBG are required to complete three rotations during their first year in the program.  Rotations provide an opportunity to explore areas for possible Ph.D. thesis research.  In addition, both students and faculty are able to test possible working relationships.

During the beginning of the Fall semester, MBG students attend Rotation Talks in which faculty who are actively seeking graduate students discuss their research.  All first-year students are expected to attend all of these talks.  

To arrange for laboratory rotations, students should discuss the possibility of a rotation with individual faculty and arrange to rotate in the interested lab at a mutually agreed-upon time.  At the end of the rotation talk period, students should have decided on their first rotation.  Each rotation should be ~8 weeks in length and all should be carried out in the lab of a MBG field faculty member.  Timing of rotations is as follows:  

BMCB and GGD:

• First Rotation — 9/3/24 – 10/25/24
• Second Rotation — 10/28/24 - 12/20/24
• Third Rotation — 1/6/25 - 2/28/25
• Chair Selection Deadline — March 9, 2025

Biophysics:

• First Rotation —  8/26/24 - 9/27/24
• Second Rotation — 9/30/24 - 11/1/24
• Third Rotation — 11/4/24 - 12/6/24
• Chair Selection Deadline —  December 13, 2024

Detailed information about 2024-25 rotations can found here . (Cornell-user access only)

Please note that graduate students do not follow the undergraduate academic calendar.  Because you are paid a twelve-month stipend, you are expected to be active in academics and/or research unless the university itself is closed.  Time away during your first year should be discussed with the DGS and your rotation supervisors.  Please refer to the section on Current Student Funding for details of time away.

While no one objects to a graduate student completing a project and writing a paper for publication during a rotation, no one expects it either!  What is expected is self-motivated earnest effort, independent thinking, and the fullest participation possible in the intellectual life of the laboratory, culminating in a written description of the project and record of the progress made.  You should have a frank discussion of lab expectations at the beginning of each rotation and again at the end for an evaluation and constructive feedback on the rotation.  Regular communication with your faculty supervisor(s), and, in subsequent years, your Special Committee, is a vital component of your success.  

Usually, by the end of the third rotation, you will have had a conversation with faculty whose lab you are interested in joining for your thesis research.  Please note, faculty should not commit to accepting a student into their lab until the end of the third rotation.  This is designed to ensure all students have the fair chance of completing their third rotation before faculty make their final decisions.  However, students are encouraged to have a clear and honest discussion with interested faculty about the possibility of joining their labs ahead of time, in order to gauge the likelihood of joining a particular lab, and whether an additional rotation will be necessary.  

All first-year students are expected to have been accepted into a lab by December 13 (Biophysics) and March 9 (BMCB/GGD).  You must inform the DGS and GFAs by that date; there is administrative work that is required to shift your funding and we do not want you to experience a gap in your stipend. In order to make your choice official in the University system, please go to your Student Center , remove the DGS of your Field as your temporary Chair (“DGS” role) and add your faculty mentor as Chair.

Students who need to initiate the fourth rotation should consult with the DGS. A student who is unable to identify a mentor by December 13 (Biophysics) and March 9 (BMCB/GGD) will not be able to continue in the program.

The evaluation of first-year students is based on grades received from courses taken and rotation evaluations.  Students should refer to the Benchmarks document received at Orientation (and available in the MBG Graduate Community Intranet ) for details of courses to be taken in the first year.  Students are expected to take a minimum of 4.5 credits of GRADED courses each semester during their 1st year in the program. Rotation Evaluations :  Supervising faculty in each rotation are required to meet with the student to discuss rotation expectations before the start of the rotation and performance at the end.  Constructive feedback is important for students and should be designed to help them not only in subsequent rotations but also in their graduate program.  A Rotation Evaluation Form must be filled out by the faculty and the signed copy given to the GRA.  

These evaluations are vital to the evaluation of the first-year class.  Advisory Committees from each field meet at the end of each year to evaluate the first year class. Their recommendations will be discussed by the entire Field faculty at the annual field meeting. A student who has not joined a lab or is judged not to have made satisfactory progress is asked to leave the program.  

GGD/BMCB In the absence of persuasive mitigating circumstances, students with the following performance in their first two semesters will be asked to leave the program:  

• Inability to identify a thesis mentor
• OR Two C grades in core courses
• OR One C grade in core courses AND one failed rotation (grade C or below)
• OR Cumulative GPA < 3.0 in core courses AND one failed rotation (grade C or below)

Biophysics In the absence of persuasive mitigating circumstances, students with the following performance in their first semester will be asked to leave the program:  

• Two or more failed rotations (grade C or below)
• OR Two C grades in core courses AND one failed rotation (grade C or below)

• Skip to Content
• Skip to Main Navigation
• Skip to Search

Indiana University Bloomington Indiana University Bloomington IU Bloomington

The College of Arts & Sciences

• Department of Biology
• Student Portal
• News & Events

IU Biology Graduate Student’s Image Featured on the Cover of Cold Spring Harbor Protocols

Tuesday, September 10, 2024

Indiana University Bloomington graduate student Shefali Shefali was honored by having her work selected for the cover of the September issue of Cold Spring Harbor Protocols . Shefali is a Ph.D. candidate in the Genome, Cell, and Developmental Biology (GCDB) program in the Department of Biology, conducting research in the Tennessen lab . Cold Spring Harbor Protocols is an interdisciplinary journal that provides a definitive source of research methods in cell, developmental and molecular biology, genetics, bioinformatics, protein science, computational biology, immunology, neuroscience and imaging. 

In this Q&A, Shefali discusses her research, the cover image and more. 

Q: How would you caption the image on the cover? I always call it “The Colorful Brain.”

The image shows the central nervous system (CNS) from third-instar Drosophila larvae (juvenile brain), highlighting the neural stem cells in red and ganglion mother cells in green. The Drosophila larval brain has a unique structure of having two optic lobes and a ventral nerve cord - and in this image one can appreciate the heterogenous cellular morphology of the central brain and optic lobes. Many studies have shown that systemic metabolism can have chronic impacts on brain development, but my study would be showing a novel mechanism by which glucose dependent signals can non-autonomously regulate cellular homeostasis within the brain. My work will be important to understand how expression patterns of metabolic enzymes, which are often thought of as housekeeping enzymes not only maintain systemic energy homeostasis, but also significantly impact cell fate decisions to have wide-range health implications. My work has been appreciated in my field, and I was recently selected to present a platform talk in the “Disease Models and Aging" session at The Allied Genetics Conference (TAGC), 2024. Moreover, my work has also been recognized at IU and I have been awarded with 2023-24 IUCOAS Kindig fellowship, 2024 Briggs fellowship and 2024 Dona Graam fellowship to support my graduate training.

Q: How was it selected, and what was your reaction? The CSHL press executive editor, Dr. Alejandro Montenegro-Montero had made an announcement for a call for Drosophila brain images, and I sent mine. I was ecstatic and beyond thrilled to find out that my image had been chosen.

Q: Is there anything else you’d like to add? I serve as the elected graduate student representative at the flyboard, an international Drosophila advisory board, that oversees the international Drosophila community. In this role, I organized a “Behind the Science Stories” workshop at The Allied Genome Research Conference 2024 , which featured some of the most incredible scientists in the country. The workshop was very well received by both early career faculty members and trainees – as they learned what goes into running a lab, finishing a story and fighting unseen battles.  Additionally, I am working on establishing a mentor-mentee program for the early career scientists in the fly community.

• Faculty + Staff Intranet

Department of Biology social media channels

• College of Arts & Sciences

• Biology B.S.
• Biology B.A.
• Special B.S. for Pre-Professional Students
• Biology Minor
• Biotechnology B.S.
• Biotechnology B.A.
• Biotechnology B.S./M.S.
• Biotechnology Minor
• Microbiology B.S.
• Microbiology B.A.
• Microbiology Minor
• Molecular Life Sciences
• Biology Exemption Exams
• Course Scheduling
• Course Evaluations
• Biology X490 Independent Study
• Research Opportunities
• Teaching Assistants
• Eligibility
• Honors Courses
• Honors Thesis
• Honors Faculty Advisors
• How to Apply
• Internships
• Evolution, Ecology, and Behavior Ph.D.
• Genome, Cell, and Developmental Biology Ph.D.
• Microbiology Ph.D.
• Biotechnology M.S.
• Multidisciplinary and Affiliated Programs
• Transfer credits
• Awards + funding
• Financial Support
• Teaching Support
• Dissertation & Thesis Support
• Services & Outreach
• Submit News to Biology
• Computing Services
• Constant Temperature Rooms
• Lactation Room
• Biology Bldg. 123
• Biology Bldg. 248
• Biology Bldg. A310
• Biology Bldg. 422
• Biology Bldg. 510
• Myers Hall 115
• Myers Hall 140
• Myers Hall 209
• Myers Hall 311
• Student Academic Appointments
• Bias incident reporting
• Title IX incident reporting
• Ombudsperson

Stritch School of Medicine

Master's in Integrative Cell Biology

Understand the biology of the cell at the molecular, multicellular, tissue, and organ structure levels.

Knowledge in the Biomedical Sciences has been growing exponentially over the last decade. As we learn more about cells at the molecular level, we also discover the need to understand their interactions and functions better. Join the next generation of Biomedical researchers with a Master of Science (MS) in Integrative Cell Biology at Loyola University Chicago.

Throughout our two-year program, you will acquire a solid foundation and understanding of cell function at molecular, multicellular tissue, and organ structure levels and gain a competitive edge in pursuing a PhD, MD, or research career.

Our Commitment To You

Graduates of Loyola's MS in Integrative Cell Biology will gain the following knowledge, skills, and professional values to pursue a PhD, apply to medical school, or pursue research in academia, industry, and/or biotech.

KNOWLEDGE :

• Develop a fundamental understanding of cellular functions in various disciplines such as cancer, neuroscience, and immunology

SKILLS : 

• Gain critical-thinking skills to help you solve biological problems
• Understand molecular and cellular technologies through your lab work

PROFESSIONAL VALUES :

• Incorporate and apply Ignatian values into your work
• Strengthen your presentation skills
• Professionally interact with peers

Students complete one year of coursework, followed by a second year focusing on research and developing their thesis. In addition to their first-year coursework, students will register for journal club and a seminar series, and select up to three laboratories to complete their rotations.  Students must complete a minimum of 30 credits during this two year MS program.

First-year coursework: 

SEMESTER 1 

• Bioethics in Biomedical Science
• Cell Biology
• Methods in Biomedical Science
• Mollecular and Cellular Biochemistry
• Rotations in up to three labs

SEMESTER 2 

• Biostatistics
• An elective course, such as Immunology, Neuroscience, Signal Transduction, or Molecular Oncology, depending on your thesis focus

Through the three rotations in the first and second semesters, students will meet with the GPD and a potential mentor to select a laboratory where they will conduct research. Students will receive their degree upon successful defense of their thesis.

Course Catalog

• Integrative Cell Biology MS Course Catalog

Ready to apply? This is a good place to start.

DEADLINES  

We accept applications for the MS in Integrative Cell Biology on a rolling basis. Early application is encouraged; we begin evaluating applications in early December. The deadline for all application materials is June 15.   Please note the only documents that cannot be uploaded with an application are transcripts and offical test scores.  Please email those documents directly to: [email protected]  

APPLICATION PROCESS  

1. COMPLETED APPLICATION

• Apply online  (preferred option) or mail application to: Graduate and Professional Enrollment Management Loyola University Chicago 820 N. Michigan Avenue, Suite 1200 Chicago, IL 60611

2. OFFICIAL TRANSCRIPTS

• Applicants should have completed at least a bachelor's degree to apply. Transcripts for all undergraduate and graduate work are required for admission.

3. LETTERS OF RECOMMENDATION

• Applicants must submit three letters of recommendation.

4. STATEMENT OF PURPOSE

• Your statement of purpose should be a brief, one page statement that explains your interest in this program.

5. INTERVIEW

• An virtual interview is required for admission. International applicants can interview by phone or video conference.

INTERNATIONAL APPLICANTS

International applicants must have a degree equivalent to a U.S. Bachelor's degree and are required to submit the above documents as well as:

• TOEFL or IELTS scores
• A Declaration and Certification of Finances Form
• Evaluations of international transcripts sent by any member organization of  NACES (National Association of Credential Evaluation Services) 

Please visit our  International Student Requirements  page for more details!

Questions? Contact Student Program Recruiter Patrick Hulseman .

Request more information about our graduate programs

WHERE ARE YOU LOCATED?

The Integrative Cell Biology Program is located at Loyola University Chicago's Health Sciences Campus in Maywood, about 12 miles west of downtown Chicago and about one-half mile south of the Eisenhower Expressway (I-290) on First Avenue.

WHAT IS YOUR GRE CODE?

If you choose to submit your GRE score, our code is 1412.

CAN I BEGIN THE MASTER'S DEGREE PROGRAM DURING THE SPRING SEMESTER?

No. We admit students into our Master's degree program for Fall semesters only; we start our program at the beginning of August.

Tuition and Financial Aid

The Biomedical Graduate Programs and Loyola's Financial Aid Office are committed to helping students secure the financial resources to make their education at Loyola affordable.

• Financial Aid

Faculty mentor students and guide them through their research.

Jordan R. Beach, PhD Assistant Professor, Cell and Molecular Physiology Case Western Reserve University PhD, Cell Physiology, 2011 University of Mount Union BS, Biology, 2004

John Callaci, PhD Associate Professor, Orthopaedic Surgery Loyola University of Chicago, PhD Loyola University of Chicago, MS

Edward M. Campbell, PhD Assistant Professor, Department of Microbiology and Immunology University of Illinois, Champaign-Urbana BS - Life Sciences/Bioengineering, 1998 University of Illinois at Chicago PhD - Microbiology and Immunology, 2004

Valerie Chai, PhD Professor, Cancer Biology Cornell University PhD, Microbial and Eukaryotic Genetics, 1999

Mashkoor Choudry, PhD Professor, Surgery, Microbiology and Immunology Aligarh Muslim University PhD, Microbiology, 1988 Aligarh Muslim University MSc, Zoology, 1983

Mitchell Denning, PhD Associate Dean for Graduate Education, Stritch School of Medicine Professor, Cancer Biology, Molecular Pharmacology and Therapeutics University of Arizona BS, Biochemistry, 1986 University of Wisconsin/Madison PhD, Human Cancer Biology, 1991

Irida Kastrati, PhD Assistant Professor, Cancer Biology University of Illinois Chicago BA, Chemistry, 2003 University of Illinois Chicago PhD, Cancer, Womens Health, Toxicology, Estrogen Carcinogenesis, Breast Cancer, Drug Design

Peter Kekenes-Huskey, PhD Associate Professor, Cell and Molecular Physiology California Institute of Technology PhD, Chemistry, 2009 University of North Carolina Asheville BS Chemistry, 2001

Ivana Kuo, PhD Associate Professor, Cell and Molecular Physiology Australian National University PhD, Neuroscience, 2010 University of Western Australia BSc, Molecular Medicine, 2004

Phong T. Le, PhD Graduate Program Director, Integrative Cell Biology Professor, Deparment of Microbiology and Immunology University of Illinois BS, Microbiology, 1980 The Ohio State University MS, Immunology, 1982 PhD, Immunology, 1985

Michael Nishimura, PhD Professor, Surgery University of Maryland Baltimore County  PhD, Biological Sciences, 1989 University of Maryland Baltimore County MS, Biological Sciences, 1984

Patrick Oakes, PhD Associate Professor, Cell and Molecular Physiology Brown University MS, Physics, 2005 Brown University PhD, Physics 2009

Clodia Osipo, PhD Associate Professor, Cancer Biology Loyola University Chicago PhD, Molecular and Cellular Biochemistry, 2002

Toni R. Pak, PhD James R. DePauw Professor and Chair, Department of Cell and Molecular Physiology University of Colorado at Boulder BA, Environmental, Population and Organismic Biology, 1995 MA, Education (Science), 1997 PhD, Neuroscience, 2002

Karie Scrogin, PhD Professor, Molecular Pharmacology and Neuroscience Oregon Health Sciences University PhD, Behavioral Neuroscience, 1991 Oregon Health Sciences University MS, Behavioral Neuroscience

Susan Uprichard, PhD Professor Medicine and Microbiology/Immunology Harvard University PhD, Virology, 1996

Ali Vaziri-Gohar, PhD Assistant Professor, Cancer Biology Kerman University of Medical Sciences MS, Biochemistry, 2006 New Mexico State University  PhD, Molecular Biology, 2015

Derek Wainright, PhD Associate Professor, Cancer Biology Loyola University Chicago PhD Cell Biology, Neurobiology, and Anatomy, 2009 Loyola University Chicago MS, Cell Biology, Neurobiology, and Anatomy, 2004

Qun Sophia Zang, PhD Associate Professor, Surgery University of Science and Technology of China BS, Molecular Biology, 1990 The City University of New York PhD, Biology/Biological Sciences, 1996

The program prepares students for a career in research laboratories in the Biomedical industry, Pharmaceutical industry or as primary or secondary educators. Students graduating with an MS are competent and competitive if they pursue a PhD or MD.

• Accessibility Options:
• Skip to Content
• Skip to Search
• Skip to footer
• Office of Disability Services
• Request Assistance
• 305-284-2374
• High Contrast
• School of Architecture
• College of Arts and Sciences
• Miami Herbert Business School
• School of Communication
• School of Education and Human Development
• College of Engineering
• School of Law
• Rosenstiel School of Marine, Atmospheric, and Earth Science
• Miller School of Medicine
• Frost School of Music
• School of Nursing and Health Studies
• The Graduate School
• Division of Continuing and International Education
• People Search
• Class Search
• IT Help and Support
• Privacy Statement
• Student Life

• Search Site
• Main College
• College News

Biomedical Engineering

• Chemical, Environmental and Materials Engineering
• Civil and Architectural Engineering
• Electrical and Computing Engineering
• Industrial Engineering
• Mechanical and Aerospace Engineering
• Letter from the Chair
• Department Clusters
• Undergraduate
• Imaging, Optics, & Lasers
• Biomechanics, Microfluidics, Biomaterials and Tissue
• Neural Engineering, Signals, & Instrumentation
• News and Events
• Advisory Board
• PhD Students
• Certificate Program
• Medical Physics Program
• Student Publications
• History and Current Enrollment

PhD in Medical Physics

• Listening Exercise
• Communications
• graduate programs

PhD in Medical Physics | College of Engineering | University of Miami

The PhD in Medical Physics Program focuses on training students’ research ability and experience in the field of medical physics with an emphasis on radiation therapy, in addition to the course work required by the MS in Biomedical Engineering – Medical Physics Program. Students graduating from the program are required to take the American Board of Radiology (ABR) exam and to apply for medical physics residency programs. Students are encouraged to seek academic positions after graduating from the program.

Students will complete required coursework by the program and will join research projects in the Department of Radiation Oncology, or other collaborative departments or clinical sites. PhD students in the program will take two qualify exams. The first one is the general qualify exam required by the Department of Biomedical Engineering, usually after two-semester study and before the third semester starts. The second qualify exam is required by the Medical Physics Graduate Program, usually after all coursework has been completed.

The Medical Physics curriculum is designed to provide students with the technical and intellectual skills required for successful careers in the field of medical physics. In addition to the coursework required by the Biomedical Engineering PhD program, PhD students enrolled in the medical physics program must successfully complete 32 medical physics course credits, at least 12 credits in research dissertation (BME 830/840) in the field of medical physics, and other requirements by the BME PhD program.  Students who received MS in Medical Physics degree from other CAMPEP-accredited programs can transfer the medical physics coursework credits.  

• 1251 Memorial Drive McArthur Engineering Building Coral Gables , FL 33146
• 305-284-2445 305-284-2445
• Academic Calendar
• Alumni & Friends
• Medical Center
• Hurricane Sports
• Parking & Transportation
• social-facebook
• social-twitter
• social-youtube
• social-instagram
• social-linkedin

Copyright: 2024 University of Miami. All Rights Reserved. Emergency Information Privacy Statement & Legal Notices

Individuals with disabilities who experience any technology-based barriers accessing the University’s websites or services can visit the Office of Workplace Equity and Inclusion .