Doctor of Philosophy (PhD) in Biology

The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of full-time study.

A summary of Biology PhD student expectations by year can be found here . Full details can be found in the Graduate Program Guide .

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program.

How to Apply    Frequently Asked Questions

Learning Outcomes

1. Demonstrate academic mastery in one of three areas of Biology: Ecology, Evolution, Behavior & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

2. Attain research expertise , including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

3. Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior, Evolution & Marine Biology; Neurobiology; or Cellular & Molecular Biology.

4. Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Coursework Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits.  See the  Graduate School of Arts & Sciences (GRS) Transfer of Credits policy  for more details. 

  • 2 semesters of Progress in Research Seminars (2 credits each):

Cell & Molecular Biology:  BI 583 & BI 584 (CM section)

Ecology, Behavior, Evolution & Marine Biology:  BI 579 & BI 580

Neurobiology:   BI 583 & BI 584 (NEURO section)

  • 1 semester grant-writing course (2 credits):

Cell & Molecular Biology:  BI 581

Ecology, Behavior, Evolution & Marine Biology: BI 671

Neurobiology: BI 581

  • 1 semester pedagogy course (1 credit): All Biology PhD students take  BI 697
  • 1 quantitative course (3–4 credits): from a list of recommended courses .
  • Additional courses vary by specific track below:

Cell & Molecular Biology

1. GRS BI 791/GRS BI 792 Graduate Rotation Credits (2 credits each/4 credits total)

2. GRS BI 753 Advanced Molecular Biology  (4 credits)

3. GRS MB 721 Graduate Biochemistry  (4 credits)

4. GRS BI 735 Advanced Cell Biology  (4 credits)

5. Two electives (8 credits, 500-level and above), see the Graduate Program Guide for recommendations

6. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology

1. Six electives  (24 credits, 500-level and above), see the Graduate Program Guide for recommendations

2. Research credits  (remaining credits)

Ecology, Behavior, Evolution & Marine Biology PhD candidate coursework is highly variable. Students, in consultation with advisors, develop a plan of coursework and research. Students are required to take a minimum of 32 credits of coursework. The remainder of the credits should be research.

Neurobiology

2. GRS BI 755 Cellular and Systems Neuroscience  (4 credits)

3. GRS BI 741 Neural Systems: Functional Circuit Analysis  (4 credits)

4. Four electives  (16 credits, 500-level and above), see the Graduate Program Guide for recommendations

5. Research credits  (remaining credits)

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge . The course provides guidance and training on pedagogy and other aspects of graduate school.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the  Cell & Molecular Biology and Ecology, Behavior, Evolution & Marine Biology curricula , this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation & Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the  GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Graduation Timeline

Forms and additional information about PhD graduation can be found on the GRS website .

9 – 12 months before proposed graduation date

  • Dissertation Prospectus & approval form due to Graduate Program Specialist for departmental review and submission to GRS

Semester prior to your intended graduation cycle

  • Intent to Graduate Form completed online

About 2 months before dissertation defense

  • Meet with Graduate Program Specialist, Director of Graduate Studies, and Faculty Advisor to review requirements for defense
  • Arrange for Special Service Appointments if committee members are not BU faculty
  • Send first draft of dissertation to readers

Once defense date is confirmed with committee

  • Reserve room(s) for public seminar and defense

At least three weeks prior to dissertation defense

  • Schedule of Final Oral Exam with Abstract Approval due to GRS
  • Properly formatted draft of dissertation submitted as PDF to [email protected]

At least two weeks prior to dissertation defense

  • Send dissertation to all committee members

At least one week prior to dissertation defense

  • Send program information to Graduate Program Specialist

See the Graduate Program Guide for final dates to submit dissertation to ETD

  • Submit final dissertation to ETD (online submission)
  • Complete electronic signature page and forward confirmation to Graduate Program Specialist along with a copy of your submitted dissertation

MS Degree (En Route to PhD)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam), and has completed 32 credits of graduate-level coursework (not including research) may apply to the Graduate School for an MS degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two: A PhD student who has taken, but has not advanced to candidacy based on the PhD qualifying examination, may still receive an MS degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for a MS degree, and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Interdisciplinary Study Options

Biology PhD students have the option to participate in the Boston University Graduate Program in Urban Biogeoscience and Environmental Health (BU URBAN), the National Science Foundation Research Traineeship Program Understanding the Brain: Neurophotonics (NSF NRT UtB: Neurophotonics), and the Biogeoscience Advanced Graduate Certificate Program . These programs require separate applications in addition to the standard Biology PhD application; those interested in BU URBAN are encouraged pre-apply .

Time Limits

Officially, the PhD must be completed within seven years after the first registration for doctoral study. PhD degrees are conferred in either May, August, or January, as specified on the GRS website . In addition, the PhD candidacy expires after the fifth anniversary of passing the Qualifying Examination. Petitions to extend this deadline are possible at the discretion of the Director of Graduate Studies, the Department Chair, and the Dean of the Graduate School, and can be obtained from the Office of the Graduate School of Arts & Sciences.

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program. PhD students are encouraged to apply for fellowships and grants at funding agencies. All domestic students should apply for NSF Graduate Research Fellowships in the Fall semester of their first or second year. 

Travel Grants may be available to assist students in their travel to professional scientific meetings; students presenting papers or posters on their research will receive first consideration.

Common Types of Funding:

Dean’s Fellowships: These are non-service fellowships allocated to first-year PhD students that do not have immediate teaching requirements.

Teaching Fellowships: These provide a stipend plus full tuition and fees for up to four full courses per semester plus a 2-credit teaching course. Teaching responsibilities usually require approximately 20 hours per week. Full or partial awards may be given.

Doctoral Research Fellowships: These awards are given to students who assist individual faculty with specific areas of research. These Research Fellowships provide a stipend and full tuition. The supervising faculty member determines the specific duties of the Research Fellow.

In addition to the above funding sources, several competitive Department awards and fellowships are available to graduate students in the Department of Biology.

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PhD Graduate Education at Northeastern University logo

The PhD program in Biology provides both a broad background knowledge base and an in-depth study of a specialized area of biology. Students can choose to pursue a concentration in Cell and Molecular Biology or Molecular Microbiology.

The PhD program in Biology emphasizes close interaction between graduate students and faculty in developing the intellectual and experimental skills required for creative independent research. To be considered for admission to the doctoral program, a student must have a bachelor’s or master’s degree, preferably in a biological science. Individuals with degrees in other natural sciences may be considered, depending on their interests and background.

A minimum GPA of 3.000 for all graduate work is required for the award of the PhD degree. All regulations of the Graduate School regarding maintenance of academic standing apply.

The Cell, Developmental, and Regenerative Biology Group studies questions related to organismal development, aging, regeneration and repair. Our unifying theme is an interest in discovering how organisms build and rebuild tissues during development, after injury, and during aging. Studies in worm, fish, axolotl, mouse, and human tissue culture are providing insights into the development, function, and aging of multiple organ systems. We use genomic and proteomic analyses, and biological imaging approaches, to understand the roles and regulatory networks underlying tissue development and repair.

Biomechanics, Neurobiology, and Behavior Group pursues distinct interests, with a special emphasis on motor learning and control, biomechanics, circadian rhythms, neurogenesis and neural regeneration. Animal models run the gamut from jellyfish and termites to fish and human subjects. Research threads include the social behavior of insects, the neural architectures underlying rhythms, movement and goal-directed behaviors in humans and model organisms, and adult neurogenesis and neuronal regeneration in teleost fish.

Molecular Microbiology Group is interested in understanding the rise of multidrug resistant pathogens. Hence the focus of this group intersects strongly with that of the University in the areas of health and security. The Molecular Microbiology group investigates the bases of dormant persister cells, uncultured bacteria, composition and function of the human and environmental microbiomes, mechanisms underlying mutagenesis, regulation of the DNA damage response, and bacterial development including biofilm formation. Further strengths include the development of platforms for the discovery of new antimicrobials.

  • Concentrations in Cell and Molecular Biology or Molecular Microbiology
  • Bachelor’s and Advanced-degree entry are possible
  • Boston residence requirement of one year of full-time graduate work on campus

Our graduates pursue careers within academia and beyond.

  • Booz Allen Hamilton
  • Albert Einstein College of Medicine
  • Agios Pharmaceuticals
  • Masschusetts General Hospital

Application Materials

Application.

  • Application fee – US $100
  • Unofficial transcripts for all institutions attended (Official transcripts required upon acceptance of admission offer)
  • Personal statement
  • Three letters of recommendation
  • GRE General – recommended, but not required
  • Proof of English Proficiency for all applicants

Our priority deadline is December 1 with rolling admissions until March 15, based on space available.

We only admit degree program students for fall semesters.

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Phd program.

Graduate students in the Department of Biology at the University of Washington receive a commitment from the entire department to support their education and research.  In addition to providing five years of assistantships, we support our many of our students through endowed departmental grants for research and travel.  Our students make excellent use of the many opportunities available to them, as evidenced by the extraordinary number of fellowships and awards received, the successful publication of high impact papers, and their regular contributions at national and international meetings.  Our students receive additional training by participating in departmental governance, curriculum development, faculty recruitment and other activities. Together these factors combine to promote an unusual degree of professionalism and community spirit.

Member of the Team

By joining the Biology Department at the University of Washington, you will become a part of a world-class research institution.  Here, graduate students are treated as colleagues by a committed faculty who work to mentor students, to provide opportunities to develop research interests, to network both on and off campus, and to publish and present their own research.

You will find a collaborative environment at the University of Washington, with faculty engaging in research with the lab next door, researchers across campus, and teams halfway across the world.  Our graduate students are encouraged to collaborate similarly, seeking additional mentors and colleagues throughout the world who compliment their research interests.

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Ph.D. in Biology

General info.

  • Faculty working with students: 50
  • Students: 91
  • Students receiving Financial Aid: 100%
  • Part time study available: No
  • Application Terms: Fall
  • Application Deadline: November 30

Amy Schmid Director of Graduate Studies Department of Biology Duke University Box 90338 Durham, NC 27708-0338

Phone: (919) 684-3649

Email: [email protected]

Website:  http://www.biology.duke.edu/

Program Description

The Department of Biology offers a diverse and integrative graduate program for students seeking a Ph.D. in the biological sciences. The department provides exciting and interdisciplinary training opportunities in the following broad range of research areas:

  • Ecology (including behavioral ecology, population biology, community ecology, physiological ecology, ecosystem analysis, and biogeochemistry)
  • Cell and molecular biology (including cellular physiology, molecular genetics, developmental biology, developmental genetics, genomics, and bioinformatics, in a variety of model systems including plants, animals, and microbes)
  • Functional organismal biology (including comparative physiology, functional morphology, biomechanics, and animal behavior)
  • Evolutionary biology (including population genetics, evolutionary ecology, evolutionary morphology, macroevolution, biogeography, and systematics)

Faculty also participate in the university programs in Cell and Molecular Biology, Computational Biology and Bioinformatics, Developmental and Stem Cell Biology, Genetics and Genomics, Ecology, and Structural Biology and Biophysics. These programs link researchers across different departments and schools at Duke, most notably the basic sciences departments in the Duke University Medical Center and the Nicholas School of the Environment and Earth Sciences. There are also strong links with the Institute of Statistics and Decision Sciences, the Institute for Genome Sciences and Policy, the Center for Nonlinear and Complex Systems, and the departments of Evolutionary Anthropology, Philosophy, and Psychological and Brain Sciences.

  • Biology: PhD Admissions and Enrollment Statistics
  • Biology: PhD Completion Rate Statistics
  • Biology: PhD Time to Degree Statistics
  • Biology: PhD Career Outcomes Statistics

Application Information

Application Terms Available:  Fall

Application Deadline:  November 30

Graduate School Application Requirements See the Application Instructions page for important details about each Graduate School requirement.

  • Transcripts: Unofficial transcripts required with application submission; official transcripts required upon admission
  • Letters of Recommendation: 3 Required
  • Statement of Purpose: Required (see departmental guidance below)
  • Résumé: Required
  • GRE General: Optional
  • GRE Subject - Biology: Optional
  • English Language Exam: TOEFL, IELTS, or Duolingo English Test required* for applicants whose first language is not English *test waiver may apply for some applicants
  • GPA: Undergraduate GPA calculated on 4.0 scale required

Department-Specific Application Requirements (submitted through online application)

Statement of Purpose Guidelines Tell us about your previous research experience, work ethic and perseverance toward goals, your preparedness and motivation for graduate study, your academic plans, including some of the research questions you would like to address in your graduate studies and how you would approach them, as well as your future career aspirations. In your statement, be sure to mention which faculty members you are interested in working with, as common interests are considered, and tell us why you believe the Duke Biology Ph.D. program is a good fit for you.

You are encouraged to include examples of educational, cultural, or other life opportunities or challenges you have experienced, and how these are likely to contribute to your overall success in our graduate program and beyond.

Writing Sample None required

We strongly encourage you to review additional department-specific application guidance from the program to which you are applying: Departmental Application Guidance

List of Graduate School Programs and Degrees

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Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2023 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

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NYU Biology’s PhD program offers training in a broad range of biological research fields, including developmental genetics, genomics and systems biology, molecular and cellular biology, evolutionary biology, and infectious disease. Our dynamic and diverse community of faculty and graduate students engages closely on all aspects of scientific investigation. Strong mentoring relationships throughout the PhD program develop students’ abilities to independently design and perform research, write grant proposals, and communicate their findings effectively to a wide audience. The combination of cutting edge research, world class faculty, and a dedication to mentoring and career building skills for our PhD students defines the PhD program at NYU Biology.

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PhD Program Overview page. View the broad range of research and training opportunities at NYU Biology.

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Ph.D. Program

Current Students

The philosophy of the PhD program, along with the Affiliated Ph.D.  Program with the Salk Institute for Biological Studies, is to provide world-class research training in the basic biological sciences to equip a diverse group of trainees for a variety of scientific careers ranging from academia and industry to education, communication, or policy. Core principles of the program are to be student centered and attuned to the goals of the trainee.

The core curriculum focuses on development of core competencies and transferable skills in critical thinking, communication, and leadership. The first year prepares students for the core of the PhD program, the development of intellectual independence and creativity through original thesis research, guided by a thesis advisor and committee. Students have a high degree of flexibility in choice of thesis advisor through the rotation program. Throughout the program, there is strong emphasis on engaged mentoring through regular committee meetings, annual reports and Individual Development Plans.

As a central hub of the thriving San Diego biosciences community, the program maintains strong partnerships with other campus units and programs through joint faculty appointments, organized research units, and research collaborations, enabling a wide range of interdisciplinary opportunities . The mission is to conduct leading edge research in the basic biological sciences. Major areas of emphasis currently include structural biology, cell biology, developmental biology, neurobiology, immunology, microbiology, virology, plant biology, ecology, and evolutionary biology. Research in the School has emphasized studies using model organisms or in vitro mechanistic approaches, with human studies and clinical research concentrated in other departments or in the Health Sciences. Current and future areas of growth include quantitative biology, data science, and the biological consequences of climate change.

  • Rigor, reproducibility, and responsibility as hallmarks of high-quality science
  • Commitment to quality mentorships, student mental health, and well-being
  • Equity, Diversity and Inclusion as integral to program admissions and retention
  • Open science practices valuing multiple research outputs and holistic assessment of scholarly excellence
  • Communication and outreach as key aspects of scientific training

As a doctoral program embedded in a large undergraduate instructional unit, our approach incorporates substantial training in teaching methodology and best practices. Our philosophy remains that teaching and research are interdependent facets of engaged scholarship.

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Orange Alert

Ph.d. in biology.

The Biology department is committed to graduate training of the highest quality. Its programs are sufficiently flexible to permit training in diverse research areas and to allow for the pursuit of interdisciplinary research programs.

Our graduate programs—leading to either the M.S. or Ph.D. degree—are offered in a wide variety of sub-disciplines within the following primary research areas: cell biology, biochemistry, molecular biology, developmental biology, ecology and plant biology, evolution, animal behavior, genetics, genomics, epigenetics, and neuroscience.

The Biology department graduate program currently averages about 50 students. About 75 percent of the students enroll directly following their undergraduate work; others come with a master's degree earned elsewhere. Virtually all departmental graduate students are supported by full tuition scholarships, as well as teaching assistantships, research assistantships, or fellowships.

APPLICATION DEADLINE : applications received on or before JANUARY 4th receive priority consideration

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Biology Graduate Programs in America

1-25 of 396 results

MIT School of Science

Cambridge, MA •

Massachusetts Institute of Technology •

Graduate School

Massachusetts Institute of Technology ,

Graduate School ,

CAMBRIDGE, MA ,

Stanford University Department of Humanities and Sciences

Stanford, CA •

Stanford University •

Stanford University ,

STANFORD, CA ,

Stanford University School of Medicine

  • • Rating 5 out of 5   3 reviews

Doctoral Student: I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. ... Read 3 reviews

3 Niche users give it an average review of 5 stars.

Featured Review: Doctoral Student says I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. .

Read 3 reviews.

Mississippi College

Graduate School •

CLINTON, MS

  • • Rating 4.37 out of 5   35

College of Science and Engineering - San Francisco State University

San Francisco State University •

SAN FRANCISCO, CA

Salisbury University

SALISBURY, MD

  • • Rating 4.52 out of 5   33

Harvard Graduate School of Arts and Sciences

Harvard University •

  • • Rating 4.56 out of 5   9 reviews

Other: I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student' in the Harvard Graduate School of Arts and Sciences. Through this School, I will be focusing my time at the John A. Paulson school of Engineering & Applied Sciences. Looking forward to wrapping up my final year on campus! ... Read 9 reviews

Harvard University ,

9 Niche users give it an average review of 4.6 stars.

Featured Review: Other says I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student'... .

Read 9 reviews.

Harvard Medical School

Boston, MA •

  • • Rating 4.94 out of 5   16 reviews

Other: Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try my best of the best….. ... Read 16 reviews

BOSTON, MA ,

16 Niche users give it an average review of 4.9 stars.

Featured Review: Other says Harvard is my dream school. Next year I will graduate the Mongolian National University of Medical Sciences. Then I will applying to Harvard Medical School. If I accepted by this school, I will try... .

Read 16 reviews.

Duke University School of Medicine

Durham, NC •

Duke University •

  • • Rating 4.76 out of 5   17 reviews

Doctoral Student: Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. ... Read 17 reviews

Duke University ,

DURHAM, NC ,

17 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. .

Read 17 reviews.

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Brown University Graduate School

Providence, RI •

Brown University •

Brown University ,

PROVIDENCE, RI ,

School of Arts & Sciences - University of Pennsylvania

Philadelphia, PA •

University of Pennsylvania •

University of Pennsylvania ,

PHILADELPHIA, PA ,

Weinberg College of Arts and Sciences

Evanston, IL •

Northwestern University •

Northwestern University ,

EVANSTON, IL ,

Vanderbilt University School of Medicine

Nashville, TN •

Vanderbilt University •

  • • Rating 5 out of 5   2 reviews

Graduate Student: Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. ... Read 2 reviews

Vanderbilt University ,

NASHVILLE, TN ,

2 Niche users give it an average review of 5 stars.

Featured Review: Graduate Student says Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. .

Read 2 reviews.

Washington University in St. Louis - Arts & Sciences

St. Louis, MO •

Washington University in St. Louis •

Washington University in St. Louis ,

ST. LOUIS, MO ,

Guarini School of Graduate and Advanced Studies

Hanover, NH •

Dartmouth College •

  • • Rating 5 out of 5   1 review

Alum: I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary learning. Guarini is well integrated within the College, which provides a unique world-class learning experience. Guarini went beyond my expectations and made this experience the best I had and will ever have in my life. The only thing I would like changed is having a GRAD diploma in Latin instead of English to keep up with College tradition. ... Read 1 review

Dartmouth College ,

HANOVER, NH ,

1 Niche users give it an average review of 5 stars.

Featured Review: Alum says I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary... .

Read 1 reviews.

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Division of Biology and Biological Engineering - California Institute of Technology

Pasadena, CA •

California Institute of Technology •

California Institute of Technology ,

PASADENA, CA ,

College of Science - University of Notre Dame

Notre Dame, IN •

University of Notre Dame •

University of Notre Dame ,

NOTRE DAME, IN ,

Biological Sciences Division - University of Chicago

Chicago, IL •

University of Chicago •

  • • Rating 4 out of 5   1 review

Master's Student: So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. ... Read 1 review

University of Chicago ,

CHICAGO, IL ,

1 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. .

Pritzker School of Medicine

Dornsife college of letters, arts and sciences.

Los Angeles, CA •

University of Southern California •

University of Southern California ,

LOS ANGELES, CA ,

Cornell University College of Arts & Sciences

Ithaca, NY •

Cornell University •

Cornell University ,

ITHACA, NY ,

Georgetown University School of Medicine

Nw Washington, DC •

Georgetown University •

  • • Rating 4.75 out of 5   4 reviews

Doctoral Student: Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1 and Step 2 preparation. ... Read 4 reviews

Georgetown University ,

NW WASHINGTON, DC ,

4 Niche users give it an average review of 4.8 stars.

Featured Review: Doctoral Student says Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1... .

Read 4 reviews.

School of Arts and Sciences - Tufts University

Medford, MA •

Tufts University •

  • • Rating 4 out of 5   4 reviews

Master's Student: My experience was mostly great! I felt supported by faculty and staff and became involved in student activities. It was just unfortunate that my on-campus experience was cut short due to the pandemic so I was not able to fully take advantage of all the program and graduate school had to offer. ... Read 4 reviews

Tufts University ,

MEDFORD, MA ,

4 Niche users give it an average review of 4 stars.

Featured Review: Master's Student says My experience was mostly great! I felt supported by faculty and staff and became involved in student activities. It was just unfortunate that my on-campus experience was cut short due to the pandemic... .

UCLA College of Letters and Science

University of California - Los Angeles •

  • • Rating 3 out of 5   1 review

University of California - Los Angeles ,

1 Niche users give it an average review of 3 stars.

Krieger School of Arts & Sciences

Baltimore, MD •

Johns Hopkins University •

  • • Rating 4.53 out of 5   19 reviews

Master's Student: I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student Advisor, Alexis has been extremely helpful in initiating my enrollment process and answering all of my questions in a timely matter. My intended Advanced Academic Program is the accelerated (2 semester), dual-modality, 40-credit M.S. in Biotechnology, Biodefense concentration. All of the anticipated course subjects are diverse and there's even a customizable core lab course on campus (at least until Summer 2024). I can't wait and I wish you all the best in your search for academic programs or professional certifications. ... Read 19 reviews

Johns Hopkins University ,

BALTIMORE, MD ,

19 Niche users give it an average review of 4.5 stars.

Featured Review: Master's Student says I have yet to enroll for Fall 2023 after receiving my acceptance letter due to a delay in my need-based financial aid from JHU. However the Homewood Campus in Baltimore is beautiful and my Student... .

Read 19 reviews.

Mellon College of Science

Pittsburgh, PA •

Carnegie Mellon University •

Blue checkmark.

Carnegie Mellon University ,

PITTSBURGH, PA ,

The Graduate School of Arts & Sciences - University of Virginia

Charlottesville, VA •

University of Virginia •

Alum: Very good in some areas, excellent in other areas, many academic choices available in all areas of study ... Read 1 review

University of Virginia ,

CHARLOTTESVILLE, VA ,

Featured Review: Alum says Very good in some areas, excellent in other areas, many academic choices available in all areas of study .

College of Sciences - Georgia Institute of Technology

Atlanta, GA •

Georgia Institute of Technology •

Other: Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer to newcomers is to prioritize building friendships early on. One regret I have as I reflect on my year is not connecting with more people. I initially thought I should always work independently, but that was a misstep in my academic journey. Remember, it's essential to strike a balance between learning and having fun. ... Read 1 review

Georgia Institute of Technology ,

ATLANTA, GA ,

Featured Review: Other says Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer... .

University of Texas - Rio Grande Valley College of Sciences

University of Texas - Rio Grande Valley •

EDINBURG, TX

  • • Rating 5 out of 5   2

College of Arts and Sciences - American University

American University •

WASHINGTON, DC

University of Pittsburgh

PITTSBURGH, PA

  • • Rating 4.43 out of 5   74

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Graduate Admissions

Helpful guides for applying to the biology phd program, tips for applying to the cmob track, tips for applying to the eco/evo track.

The PhD program trains students for careers in research and teaching. In addition to a wide range of courses, the PhD program offers informal but intensive instruction in laboratories and seminars, characterized by close contact between students and their faculty advisors. Biology is one of fourteen home programs  in the  Biosciences,  which span the School of Medicine and the School of Humanities and Sciences.

How to Apply

For the 2023-2024 PhD application season, application processes will be run through existing Biology (School of Humanities & Sciences)  and the Oceans Department(Stanford Doerr School of Sustainability). 

Previously part of the Biology Department , Hopkins Marine Station is now part of the Oceans Department within the Stanford Doerr School of Sustainability (SDSS).  Several faculty took appointments in Oceans, while others chose to stay in Biology; still others have joint appointments.  Additionally, several faculty members in Ecology/Evolutionary Biology now have joint appointments within Biology and SDSS. Based on these changes, the following information will provide further guidance for the application process. 

  • Faculty with 100% appointments in Biology, will only recruit through Biology.  The Biology application will be used.
  • Ecology/Evolutionary Biology faculty with 50/50 appointments in both Biology and SDSS will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Ecology/Evolutionary faculty with joint Biology/SDSS appointments, you should reach out to them directly regarding the appropriate application track to choose. 
  • 100% Oceans faculty will only be recruiting students through the Earth Systems Science application.
  • Oceans faculty appointed 50/50 in Biology and SDSS (Oceans Department), will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Oceans faculty with joint Biology/SDSS appointments, you should reach out to the individual faculty members you hope to work with regarding the appropriate application and track to choose. 

Faculty Affiliated with the Stanford Doerr School of Sustainability

Oceans Department:

Fio Micheli 100% 

Jeremy Goldbogen 100%

Giulio De Leo 100%

Larry Crowder 100%

Steve Palumbi 50% Oceans/50%Biology

Barbara Block 50% Oceans/50% Biology

*Note Chris Lowe will remain 100% in Biology

Ecology/Evolution

Liz Hadly 50% SDSS/50%Biology

Rodolfo Dirzo 50% SDSS/50%Biology

Tad Fukami 50% SDSS/50%Biology

Kabir Peay 50% SDSS/50%Biology

Chris Field 50% SDSS/50% Biology

Gretchen Daily 50% SDSS/50% Biology

Application and Deadline

Stanford Biology PhD Program applications are made through Graduate Admissions . The application deadline for Autumn Quarter 2024 matriculation is December 5, 2023 at 11:59pm pst.  The application for the Autumn 2024 cohort will be available in September 2023.  Please review the Graduate Admissions website prior to starting your application. Applications are considered for admission in Autumn Quarter only. 

Application Status

To check your application status,  click here to Visit Your Status Page .  

Requirements

  • U.S. Bachelor's degree or its foreign equivalent
  • Stanford Graduate application
  • Non-refundable application fee of $125 (or fee waiver found here ) 
  • Statement of Purpose
  • Three Letters of Recommendation (a maximum of four letters will be accepted)
  • Transcripts - Unofficial transcripts from all universities and colleges you've attended for one year or more must be uploaded to the online application.  Graduate Admissions only requires admitted applicants who accept the offer of admission to submit official transcripts that shows their degree conferral. Please do not send or have sent any official transcripts to us at this time.
  • TOEFL Test of English as a Foreign Language (required for non-native English speakers)
  • A current Curriculum Vitae/Resume

Knight-Hennessy Scholars

Join dozens of  Stanford School of Humanities and Sciences students  who gain valuable leadership skills in a multidisciplinary, multicultural community as  Knight-Hennessy Scholars  (KHS). KHS admits up to 100 select applicants each year from across Stanford’s seven graduate schools, and delivers engaging experiences that prepare them to be visionary, courageous, and collaborative leaders ready to address complex global challenges. As a scholar, you join a distinguished cohort, participate in up to three years of leadership programming, and receive full funding for up to three years of your PhD studies at Stanford. Candidates of any country may apply. KHS applicants must have earned their first undergraduate degree within the last seven years, and must apply to both a Stanford graduate program and to KHS. Stanford PhD students may also apply to KHS during their first year of PhD enrollment. If you aspire to be a leader in your field, we invite you to apply. The KHS application deadline is October 11, 2023. Learn more about  KHS admission .

Biology PhD Program General Information

Diversity, inclusion, equity, and belonging.

The Department of Biology recognizes that the Supreme Court issued a ruling in June 2023 about the consideration of certain types of demographic information as part of an admission review. All applications submitted during upcoming application cycles will be reviewed in conformance with that decision. The Department of Biology welcomes graduate applications from individuals with a broad range of life experiences, perspectives, and backgrounds who would contribute to our community of scholars. Review of applications is holistic and individualized, considering each applicant’s academic record and accomplishments, letters of recommendation, and admissions essays in order to understand how an applicant’s life experiences have shaped their past and potential contributions to their field.Please see the websites below for more resources.

  • Diversity and Engagement in Biosciences PhD Programs
  • Vice Provost for Graduate Education
  • Biology Equity and Belonging
  • Research Sites
  • Faculty Profiles and Labs
  • Inquiries about the PhD in Biology should be directed to  biologyadmissions [at] lists.stanford.edu (biologyadmissions[at]lists[dot]stanford[dot]edu)

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Department of Biology

  • Graduate Program

PhD Program

Our PhD program is designed to foster the development of scientists capable of doing advanced biological research. We offer students research opportunities in:

  • Molecular, cellular, and developmental biology
  • Evolution, ecology, genetics, and genomics

Our faculty share a common interest in genetic processes. As a result, our students benefit from the high degree of interaction and cooperation among laboratories.

Our program, taken on a full-time basis, requires a total of 90 credit hours. Credit hours include coursework, seminars, and research. The average course load is 12 credit hours per semester.

Virtual tour - Learn why our graduate program is right for you

Requirements

For a comprehensive review of the PhD requirements, see the PhD Handbook . For information about how to apply to the program and for financial support, see the application process page .

Years One and Two

Each student rotates in three laboratories in the first year, at the end of which he/she chooses a faculty advisor by mutual agreement. Students can arrange to rotate in a range of laboratories in the Department of Biology and affiliated laboratories in the University of Rochester’s Medical Center . Students also take five to six courses over their first two years (selected according to background and research interests) and serve as teaching assistants for two semesters in order to gain teaching experience.

Qualifying examinations are normally taken in June of the second year.

Completing Your Degree

After passing the qualifying examination, students form a thesis advisory committee to help guide them in their research. Completion of the PhD usually takes six years.

Communication and interaction are essential parts of being a scientist. Our students benefit from multiple opportunities to attend seminars by distinguished outside speakers, participate in formal and informal journal clubs and discussion groups, and present their research to their peers.

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Graduate program.

Duke Biology offers diverse training opportunities for students seeking a Ph.D. degree in biological sciences. Our program is highly flexible, allowing students to design the path that best fits their professional needs. Our students spend the majority of their time engaged in innovative research, and have the opportunity to interact and collaborate with researchers from a wide range of fields. 

The graduate students in the Biology Department (Biograds) are integral to the department and community. Contributions include: running weekly happy hours and prospective student recruitment weekends as well as organizing seminars, journal clubs, teaching-assistant training orientation, and diversity-themed workshops.

Program Requirements

Requirements for the Ph.D. include course work, participation in seminars relevant to your field of research, successful completion of an oral qualifying examination, one year of teaching experience, completion and defense of a research dissertation under faculty guidance.

Affiliated Graduate Programs

Duke Biology is involved in multiple University-wide graduate programs, which link researchers across different departments and schools at Duke – most notably the basic sciences departments in the  Duke University Medical Center  and the  Nicholas School of Earth and Ocean Sciences .

For Prospective Students

Learn more about our degree, our university, and our community. We'll explain the application process, financial support, and more.

  • For Current Students

We've compiled resources to assist you while working toward your Ph.D. – from requirements and fellowships, to milestones, useful Duke links, and more.

Learn more about the application process, deadlines and other resources for our candidates.

Review Application Information

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  • Exploring the Extracellular Matrix: Dr. David Sherwood
  • Cell Division's Missing Link: Dr. Masayuki Onishi
  • Listening in to Birdsong: Dr. Steve Nowicki
  • Biogeochemistry as Ecosystem Accounting: Dr. Emily Bernhardt
  • Building a Dynamic Nervous System: Dr. Pelin Volkan
  • Investigating a Key Plant Hormone: Dr. Lucia Strader
  • Imagining Visual Ecology: Dr. Sönke Johnsen
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Biological Sciences: PhD

Graduate Program in Biological Sciences

Program Handbook

The Department of Biological Sciences is the focal point for research in the life sciences at the University of Notre Dame. The doctoral program in biological sciences encompasses the broad scope of biological inquiry with several areas of study in the biomedical sciences, global health, and ecology, evolution & the environment (EEE). This uncommon level of wide-ranging yet integrative biological inquiry provides an extraordinary environment for learning, innovation and collaboration.

The program provides a wide choice of faculty advisers, many of who lead their disciplines by serving as officers of professional societies, on journal editorial boards, study sections, scientific advisory boards and as consultants for various organizations. This vibrant research community continues to expand its faculty, both within the department and with associated faculty at the Indiana University School of Medicine at South Bend adjacent to the Notre Dame campus, and other departments at Notre Dame.

Updated: 08/31/2023

  • GRE General Test optional
  • TOEFL, IELTS, or Duolingo for non-native speakers of English
  • Curriculum vitae
  • Official transcripts from each post-secondary institution; one must show conferral of a bachelor's degree. (Due upon enrollment)
  • Statement of intent
  • Three letters of recommendation
  • Unofficial transcripts from each post-secondary institution required at the time of application. (Official transcript showing conferral of a bachelor's degree due upon enrollment.)

Rebecca Wingert Director of Graduate Studies Phone: (574) 631-0907 Email: [email protected]

https://biology.nd.edu/graduate/graduate-studies-overview/

Rensselaer

  • Undergraduate

Ph.D in Biology

Our doctoral program in biology prepares students for research-oriented careers in academia, government, and the private sector. .

Candidates for the Ph.D. must satisfy the requirements of the Graduate Program Committee (GPC) including passing the first year Core Course, which serves as a qualifying exam with a grade of B or better, and passing two at least 3-credit graduate elective courses (one of which must be at the 6000 level, and one of which must be in BIOL or BCBP). Entering students must complete three rotations in the first year. A thesis advisor must be chosen from the list of participating faculty by the end of the second semester. Students must also pass a candidacy exam which consists of a written and an oral portion, and must be taken by the end of the second year of full-time study. Degree candidates also must submit a dissertation based on an original research project, present their findings in a public seminar, and defend their work in an oral examination. Additionally, all doctoral candidates are required to participate in teaching for at least one semester under the supervision of a faculty member. Seventy-two credit hours are required for the Ph.D.

Outcomes of the Graduate Curriculum

Students who successfully complete this program will be able to:

  • explain general principles and concepts from a variety of sub-disciplines within the broader field of biology.
  • demonstrate expert knowledge of a specialized field of biology and be able to ask and answer challenging questions in that field.
  • critically analyze and interpret the scientific literature and scientific presentations.
  • independently design, prepare, and execute experiments, using appropriate research techniques.
  • critically interpret research data and evaluate findings using appropriate statistical analyses.
  • independently design and execute a research strategy aimed towards answering a pressing scientific question in the field.
  • effectively communicate their scientific research and findings in a variety of written and oral formats, including the ability to prepare a manuscript of original research for publication in a peer-reviewed scientific journal.
  • discuss issues related to scientific ethics and scientific misconduct and apply ethical standards to their research and professional conduct.

Documents & Resources

2023-2024 Biological Sciences Graduate Student Handbook

PhD - Timeline of Required Forms

Department of Biology

cells

A doctoral degree is offered by the Program in Cell, Molecular, Developmental Biology, and Biophysics (CMDB) .

CMDB graduate students participate in a core curriculum including molecular biology, cellular biology, developmental biology, and biophysics. Students continue to broaden their knowledge in these areas throughout their graduate training while they specialize in their own research areas. Through this cross-training, PhDs emerge from the CMDB program prepared to tackle complex problems in 21st-century biosciences.

The CMDB program includes faculty from Johns Hopkins University’s departments of Biology, Biophysics , and Chemistry , as well as from the Carnegie Institution for Science Department of Embryology .

Other JHU Graduate Programs in Biology

  • MS in Biology
  • Part-time MS in Biotechnology
  • Part-time MS in Bioinformatics
  • Post-Baccalaureate Premedical Program

Biological Sciences

  • Mellon College of Science

Ph.D. in Biological Sciences

The mission of the Ph.D. program in Biological Sciences is to train new generations of scientific leaders who make significant and distinctive contributions to society. The department has established a broad range of resources to enable the students to define and achieve their personal mission. With this broad goal in mind, our specific objectives are to mentor students to:

Become contributing, impactful members of their discipline. This requires:

  • Being familiar with literature and history of the field
  • Understanding how the discipline has evolved and where it is headed
  • Identifying novel and important questions
  • Identifying desired personal impact on the field.
  • Mastering approaches and techniques
  • Appreciating the impact of the discipline on a broader community
  • Establishing visible profile in the field

Develop professional skills in :

  • Scientific writing (papers, proposals, traditional and social media)
  • Oral presentation
  • Team management (including interdisciplinary teams)
  • Communication, networking, connecting with community of researchers & science professionals
  • Time management, life/work balance

Establish career development plans through :

  • Establishing sense of scholarly citizenship
  • Knowing the types of careers paths following from Ph.D. training
  • Developing personal career goals
  • Appreciating skills required to achieve career goals and how to align personal training with these goals 

Ph.D. Student Research Videos

Emily simon (linstedt lab), andrew wolff (hinman lab).

  • Online Application

Beril Tutuncuoglu

" CMU provides an innovative environment, where technology meets deep scientific thinking. I received constant guidance both for my research projects and career path. I had the opportunity to explore different areas and expand my imagination about what is possible to discover in biology, by taking selective courses and consulting faculty with a broad range of expertise. Through presenting at international conferences, I established scientific collaborations, which lead to significant findings and publications, one of which is in the highly regarded journal  Nature . A key perk of being a part of the Department of Biological Sciences is the alumni network of CMU, which consists of overachieving and fearless individuals. As a recent hire of a company started by two CMU alumni, I am excited to continue my pursuit of discovering the unknown and challenging the current limits of experimental sciences ."

Santosh Chandrasekaran

" One of the characteristics of CMU that I particularly loved was the extensive levels of collaborations between research groups, even from seemingly unrelated fields. Such exquisite collaborations often further the frontiers of research and being a graduate student at CMU allowed me to lead and be a part of such high quality research. This is primarily due to the excellent faculty at CMU who are always interested in exploring new ideas and are very approachable which gave me immense encouragement  ..... Pittsburgh also provided ample opportunities to pursue my varied hobbies, be it art and music or skiing and rock climbing. This was particularly important for me as it provided a way to switch off from research and rejuvenate. "

Ken Hovis

" As a prospective student in neuroscience, most of the graduate programs I applied to were neuroscience only programs. One of the biggest reasons I am so glad I chose the Biological Sciences graduate program at CMU, is because I not only gained access to excellent training in the field of neuroscience, but I was surrounded by faculty, post-docs, and other graduate students from many other disciplines. This provided me with different perspectives on my research and enabled me to have access to, learn, and incorporate a variety of techniques from different fields during my graduate experience. It also provided me with experience working as part of an interdisciplinary team, which is a very important part of my job now as I work closely with faculty from mathematics, physics, chemistry, and biology. "

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Doctor of Philosophy - Biological Sciences

Jump to section:, learning outcomes, career possibilities, requirements.

  • Documents and Downloads

The Doctor of Philosophy (Ph.D.) in Biological Sciences within the School of Life Sciences (SoLS) consists of four sectional research concentrations that reflect the scope of modern biology: Ecology and Evolutionary Biology (EEB), Integrative Physiology (IP), Cell and Molecular Biology (CMB), and Microbiology (MB). The degree is research intensive and designed to prepare students for careers in academia, government, and industry as engaged scholars who are experts in their chosen field. Students must complete a minimum of 60 credit hours from a list of core and approved courses within their research section. They must engage in independent research that is novel and exciting culminating with a dissertation that makes an important contribution to their chosen field. As such, it is expected that their dissertation work will be published in peer reviewed journals with the student listed as first author. Successful students are also trained and expected to develop as effective teachers and educators, and each student must serve as a teaching assistant for two semesters as part of the degree program.

Available Options

Post-bachelor’s – cellular and molecular biology track, post-bachelor’s – ecology and evolutionary biology track, post-bachelor’s – integrative physiology track, post-bachelor’s – microbiology track, post-bachelor's – quantitative biology and bioinformatics track, post-master’s track, accreditation.

For information regarding accreditation at UNLV, please head over to Academic Program Accreditations .

All students graduating with a Ph.D. in Biological Sciences should be able to:

  • Master a critical set of key concepts specific for each sectional concentration.
  • Gain expertise with key methodologies and experimental techniques specific for each sectional concentration.
  • Read, comprehend, and critically evaluate the current published scientific literature.
  • Evaluate a scientific question and formulate testable hypotheses.
  • Independently design experiments to effectively test hypotheses.
  • Generate and collect reproducible data.
  • Access up-to-date methods for analyzing and tabulating data.
  • Communicate scientific results effectively in oral presentations to general and specialized audiences.
  • Communicate scientific results effectively in written reports for publication in peer-reviewed journals.
  • Apply and be competitive for extramural monies to fund research.
  • Instruct and engage students and members of the community at all levels to appreciate the importance of biology in their lives.
  • Career Possibilities 118.53 KB

Documents/Downloads

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  • BIOL 796 67.45 KB

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Graduate handbooks.

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Degree Contact

Keala "ala" kiko.

Director of Graduate Programs

School of Life Sciences

Life sciences involves studies of living organisms and their life processes, including their evolution and relationships with other living organisms and our planet. The courses and programs offered by the School of Life Sciences are designed for those students pursuing professional careers in medicine, science, and science education.

College of Sciences

The School of Life Sciences offers programs that meet the needs of students intending to enter the workforce or pursue advanced training in the sciences, medicine, and other professional and technical fields. We provide a well-rounded foundation in natural, physical, and mathematical sciences that can set students up for successful careers and professional programs.

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University of Northern Colorado

Biological Education

Doctor of Philosophy

Advance your career with a doctorate that combines research expertise with outstanding teaching ability and experience.

Biological Education PhD at UNC

  • Programs at the University of Northern Colorado
  • Biological Sciences
  • Biological Education PhD

Join a Pioneering Program for Biology Educators and Researchers

Recent graduates of UNC’s Biological Education Ph.D. program have a high job placement rate. Our program helps you stand out in the market by providing comprehensive training for positions in academia, government and the private and nonprofit sectors. In addition to offering strong opportunities for biological field and laboratory research and publication, our Ph.D. program provides extraordinary training in STEM education. You’ll graduate with an in-demand skill set that integrates research expertise with outstanding teaching ability and experience.

Established in the early 1990s, this pioneering program was among the first in the nation to emphasize the teaching of biology. You’ll get extensive hands-on experience as a teaching assistant and classroom leader, culminating in the opportunity to develop and teach your own college course. University biology and other employers place high value on this training. Many of our Ph.D. candidates have multiple job offers waiting for them at graduation.

Degree Details

Credits required:, location/delivery:.

Cost Estimator

Degree Option

Ph.d. in biological education.

UNC’s Ph.D. in Biological Education offers options you won’t find in most doctoral programs. Focus your dissertation studies on biological research or on the teaching and learning of biology, and enrich your education by working side-by-side with UNC's biology faculty who excel in both types of research.

Take the next step! Explore courses, costs and admission requirements.

Degree Requirements

Learn more about the Ph.D. Program

"The biology graduate program at UNC was a great fit for me. The PhD program provides a nice blend of science and science education. My skills as both a biologist and an educator were strengthened through my experiences. That doesn't happen in all graduate programs. I think that having a solid background in biology as well as a track record of successful teaching really helped me to stand out in the job application process. I feel I was well prepared for my immediate future as an Assistant Professor of Biology at a liberal arts college."

Your Future in Biological Education

Because of its inclusion of training in Biological Education, UNC’s Ph.D. program equips you to contribute to the future of STEM education. A broad national consensus on the need to improve outcomes in STEM education creates a broad range of employment opportunities in colleges and universities, government agencies, nonprofit organizations and the private sector.

Consider a Ph.D. in Biological Education if you:

  • Are committed to biological research
  • Love to teach
  • Seek to further the goals of national STEM education policy
  • Have an interest in expanding the frontiers of biological research

You’ll learn:

  • Research methods in the area of your choice
  • How to design and teach a college biology course
  • Scientific writing and publishing

Sample courses:

  • Advanced Genetics
  • Immunology and Disease
  • Conservation Biology
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Beyond the Classroom

Our Ph.D. candidates work as teaching assistants and all eventually design and teach an original course. In addition, many of our Ph.D. candidates gain experience as research assistants, working in fields as diverse as molecular biology, evolutionary biology, human health and biomedical science, biological pedagogy and other sub-disciplines.

Where can your degree take you?

  • Tenure-track faculty position in a university biology department
  • Conservation nonprofit or government agency
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  • Medical research institute
  • Postdoctoral positions
  • Biotechnology careers

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  2. PhD in Biology: Biology Courses

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  3. 2024 Best Online PhD in Biology Programs [Doctorate Guide]

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  4. [Infographic] Where Will a PhD in Biology Take You? A faculty job is a

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  5. 2023 Best Online PhD in Biology [Doctorate Guide]

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  6. PhD in Biology- Admission Process, Requirements, Fees, Duration, Job

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  1. PhD Admission 2024 PhD Entrance Exam Updates Shivaji University Kolhapur

  2. PhD

  3. Major in Neuroscience

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  5. Fully Funded PhD in Neuroscience at Harvard University

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COMMENTS

  1. Doctor of Philosophy (PhD) in Biology

    The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of ...

  2. Ph.D. Program

    The Biology Ph.D. program is part of the larger Biosciences community at Stanford, which includes doctorate programs in the basic science departments at Stanford Medical School. There are two tracks within the Biology Ph.D. program: Cell, Molecular and Organismal Biology. Ecology and Evolution. (Previously a part of the Department of Biology ...

  3. Top Biology Doctorate Degrees & Graduate Programs 2024+

    Biology Doctorate Degrees. The PhD in Biology degree is an in-depth, research degree and the highest level of education in the field. In most biology PhD programs, students choose a specific area in the biological sciences to anchor their studies. Some of the options might include Cell and Molecular Biology, Ecology and Evolution, Neurobiology ...

  4. Biology

    The PhD program in Biology provides both a broad background knowledge base and an in-depth study of a specialized area of biology. Students can choose to pursue a concentration in Cell and Molecular Biology or Molecular Microbiology. The PhD program in Biology emphasizes close interaction between graduate students and faculty in developing the ...

  5. PhD Program

    PhD Program. Graduate students in the Department of Biology at the University of Washington receive a commitment from the entire department to support their education and research. In addition to providing five years of assistantships, we support our many of our students through endowed departmental grants for research and travel.

  6. Ph.D. in Biology

    The Department of Biology offers a diverse and integrative graduate program for students seeking a Ph.D. in the biological sciences. The department provides exciting and interdisciplinary training opportunities in the following broad range of research areas:

  7. Biology, PhD < University of Pennsylvania

    Biology, PhD. The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student's thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology ...

  8. NYU Biology PhD Program

    NYU Biology's PhD program offers training in a broad range of biological research fields, including developmental genetics, genomics and systems biology, molecular and cellular biology, evolutionary biology, and infectious disease. Our dynamic and diverse community of faculty and graduate students engages closely on all aspects of scientific ...

  9. Ph.D. Program

    The philosophy of the PhD program, along with the Affiliated Ph.D. Program with the Salk Institute for Biological Studies, is to provide world-class research training in the basic biological sciences to equip a diverse group of trainees for a variety of scientific careers ranging from academia and industry to education, communication, or policy.

  10. Ph.D. in Biology

    Our graduate programs—leading to either the M.S. or Ph.D. degree—are offered in a wide variety of sub-disciplines within the following primary research areas: cell biology, biochemistry, molecular biology, developmental biology, ecology and plant biology, evolution, animal behavior, genetics, genomics, epigenetics, and neuroscience. The ...

  11. 2023-2024 Top Biology Graduate Programs

    My intended Advanced Academic Program is the accelerated (2 semester), dual-modality, 40-credit M.S. in Biotechnology, Biodefense concentration. All of the anticipated course subjects are diverse and there's even a customizable core lab course on campus (at least until Summer 2024).

  12. Graduate Admissions

    Stanford Biology PhD Program applications are made through Graduate Admissions. The application deadline for Autumn Quarter 2024 matriculation is December 5, 2023 at 11:59pm pst. The application for the Autumn 2024 cohort will be available in September 2023. Please review the Graduate Admissions website prior to starting your application.

  13. PhD Program : Graduate Program : Department of Biology : University of

    University of Rochester. 402 Hutchison Hall. P.O. Box 270211. Rochester, NY 14627. (585) 275-3835. Our PhD program is designed to foster the development of scientists capable of doing advanced biological research. We offer students research opportunities in:

  14. Graduate Program

    Graduate Program. Duke Biology offers diverse training opportunities for students seeking a Ph.D. degree in biological sciences. Our program is highly flexible, allowing students to design the path that best fits their professional needs. Our students spend the majority of their time engaged in innovative research, and have the opportunity to ...

  15. 308 PhD programmes in Biology in United States

    The Biological Sciences - Cell Biology and Molecular Genetics degree from Kent State University is an individualized program of study and research that prepares students for career opportunities in teaching and research in academic, governmental and industrial settings. Ph.D. / Full-time / On Campus.

  16. Best 308 Biology PhD Programmes in United States 2024

    308 Biology PhDs in United States. Molecular and Cell Biology. University of Maryland Baltimore County (UMBC) Baltimore Highlands, Maryland, United States. Biological Sciences - Cell Biology and Molecular Genetics. Kent State University. Kent, Ohio, United States. Biological and Agricultural Engineering.

  17. Biological Sciences: PhD

    The Department of Biological Sciences is the focal point for research in the life sciences at the University of Notre Dame. The doctoral program in biological sciences encompasses the broad scope of biological inquiry with several areas of study in the biomedical sciences, global health, and ecology, evolution & the environment (EEE).

  18. Biology

    Candidates for the Ph.D. must satisfy the requirements of the Graduate Program Committee (GPC) including passing the first year Core Course, which serves as a qualifying exam with a grade of B or better, and passing two at least 3-credit graduate elective courses (one of which must be at the 6000 level, and one of which must be in BIOL or BCBP).

  19. Graduate

    A doctoral degree is offered by the Program in Cell, Molecular, Developmental Biology, and Biophysics (CMDB). CMDB graduate students participate in a core curriculum including molecular biology, cellular biology, developmental biology, and biophysics. Students continue to broaden their knowledge in these areas throughout their graduate training while they specialize in their own research areas.

  20. Ph.D. in Biological Sciences

    The mission of the Ph.D. program in Biological Sciences is to train new generations of scientific leaders who make significant and distinctive contributions to society. The department has established a broad range of resources to enable the students to define and achieve their personal mission. With this broad goal in mind, our specific ...

  21. Doctor of Philosophy

    The Doctor of Philosophy (Ph.D.) in Biological Sciences within the School of Life Sciences (SoLS) consists of four sectional research concentrations that reflect the scope of modern biology: Ecology and Evolutionary Biology (EEB), Integrative Physiology (IP), Cell and Molecular Biology (CMB), and Microbiology (MB).

  22. 9 Careers After a PhD in Biology (With Salaries)

    Here are nine careers that you can pursue after earning a Ph.D. in biology. For the most up-to-date Indeed salaries, please click on the links below: 1. Postsecondary biological sciences teacher. National average salary: $53,712 per year Primary duties: Postsecondary biological sciences teachers lecture biology students on various related ...

  23. Join a Pioneering Program for Biology Educators and Researchers

    In addition to offering strong opportunities for biological field and laboratory research and publication, our Ph.D. program provides extraordinary training in STEM education. You'll graduate with an in-demand skill set that integrates research expertise with outstanding teaching ability and experience. Established in the early 1990s, this ...