phd programs marine biology

Doctoral Programs

Interested in applying to our MS or PhD programs? View presentations from this year's info sessions .

The Scripps Institution of Oceanography PhD program is organized into three separate academic programs. Each is responsible for all graduate educational activities in its area, including teaching, advising, and examining. Prospective PhD students will apply directly to one or more.

Academic programs

• Climate-Ocean-Atmosphere Program
• Geosciences of the Earth, Oceans, and Planets
• Ocean Biosciences Program

A graduate student's work normally will be concentrated in one of several curricular groups under one of the academic programs:

Climate-Ocean-Atmosphere Curricular Groups

• Applied Ocean Science
• Climate Sciences
• Physical Oceanography

Geosciences of the Earth, Oceans, and Planets Curricular Groups

• Geosciences
• Marine Chemistry and Geochemistry

Ocean Biosciences Curricular Groups

• Biological Oceanography
• Marine Biology

The interdisciplinary nature of research in marine and earth sciences is emphasized; students are encouraged to take courses from various UC San Diego departments and to consider interdisciplinary research projects.

Additional programs

The Program for Interdisciplinary Environmental Research (PIER) is a graduate specialization for select doctoral students who seek solutions to today's environmental challenges.

This PhD Specialization is designed to allow students to obtain standard training in their chosen field and an opportunity to interact with peers in different disciplines throughout the duration of their PhD training. Admission to PIER is a competitive process with 6 - 8 students granted admission each year from the ten participating UC San Diego departments.  More information can be found on the PIER Program website .

A joint graduate group from the Geophysics Program of Scripps Institution of Oceanography at the University of California San Diego (UC San Diego) and the Department of Geological Sciences at San Diego State University (SDSU) offers a Joint Doctoral Program in Geophysics.

The complementary specialties and ongoing, vigorous collaborations between the two groups result in two focus areas: earthquake science and applied geophysics. Integrating geophysics at UC San Diego and SDSU provides outstanding opportunities for students to develop the skills needed to address important local, regional, and global societal problems where geophysics can contribute to the solutions.

Strong capabilities will be in:

More information can be found on the Joint Doctoral Program website .

Graduates will be prepared to begin rewarding geophysics careers and assume leadership roles as university faculty, government scientists, and industry researchers. Joint UCSD and SDSU committees will administer and monitor the admission, advising, evaluation, graduation, and all other academic processes related to the joint doctoral program. Students will spend at least one academic year of residency at each campus.

A Doctor of Philosophy (PhD) degree in Geophysics will be awarded upon completion of the program in the names of The Regents of the University of California on behalf of UC San Diego and The Trustees of the California State University on behalf of SDSU. Prospective applicants will apply through SDSU.

The Department of Scripps Institution of Oceanography offers a concurrent degree program allowing interested PhD students to complete an MBA at the Rady School of Management .

Students who are admitted to Scripps may, with the consent of their academic advisor, apply to Rady.  Application to the MBA program is through the usual admissions process.  Students may begin the MBA program no earlier than after the completion of their departmental exam and no later than the fall quarter following their advancement to candidacy.

An extensive independent study, jointly supervised by Scripps and Rady faculty, enables the student to develop linkages between Scripps and Rady studies. Interested students are encouraged to consult early with Rady MBA Admissions and with their Scripps academic advisors.

Ph.D. in Marine Science and Conservation

General info.

• Faculty working with students: 25 (15 in Beaufort and 9 in Durham)
• Students: 38
• Students receiving Financial Aid: 100%
• Application terms: Fall
• Application deadline: December 7

James Hench Director of Graduate Studies Duke Marine Laboratory 135 Duke Marine Lab Road Beaufort, NC 28516-9721

Phone: (252) 504-7585

Email: [email protected]

• https://nicholas.duke.edu/academics/doctoral-degrees
• https://nicholas.duke.edu/marinelab

Program Description

The PhD in MSC  is designed to ensure that students receive detailed training in a particular discipline of marine natural or social science, while being exposed to sufficient interdisciplinary knowledge that they are able to synthesize information from both the social and natural sciences and apply that information to policy-making.  These students will be prepared for careers either in university teaching or research, or outside of the university involving the application of science to policy-making. 

Doctoral students in MSC emphasize research as the major part of their degree programs.  The committee, chaired by the student’s faculty advisor, will recommend which of the program’s required courses students should take along with any additional courses, and generally help to shape the student’s research program.

The bulk of a student's time will be spent at the Duke Marine Lab in Beaufort, NC. Students often spend one or two semesters taking graduate level classes on the Durham campus before moving to Beaufort to complete their research; however, residence in Durham is not a requirement.

In addition to MSC and other course work, MSC students are required to present short research update seminars in years 2-4 of their program, complete a preliminary examination, and complete a dissertation. The dissertation is presented in a public seminar and defended to the committee in a dissertation examination.

• Marine Science and Conservation: PhD Admission and Enrollment Statistics
• Marine Science and Conservation: PhD Completion Rate Statistics
• Marine Science and Conservation: PhD Time to Degree Statistics
• Marine Science and Conservation: PhD Career Outcomes Statistics

Application Information

Application Terms Available:  Fall

Application Deadline:  December 7

Graduate School Application Requirements See the Application Instructions page for important details about each Graduate School requirement.

• Transcripts: Unofficial transcripts required with application submission; official transcripts required upon admission
• Letters of Recommendation: 3 Required
• Statement of Purpose: Required
• Résumé: Required
• GRE Scores: GRE General (Optional)
• English Language Exam: TOEFL, IELTS, or Duolingo English Test required* for applicants whose first language is not English *test waiver may apply for some applicants
• GPA: Undergraduate GPA calculated on 4.0 scale required

Writing Sample None required

We strongly encourage you to review additional department-specific application guidance from the program to which you are applying: Departmental Application Guidance

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The Marine Biology and Ecology (MBE) graduate program focuses on a wide range of field, laboratory, and theoretical coursework in research areas such as coral reef studies, biological oceanography, and marine biology, fisheries biology, and the biology and behavior of marine animals. MBE Ph.D. students can choose from a diversity of research areas and coursework that provide a broad educational perspective. 

Admission Requirements

Most successful applicants have a bachelor’s degree in biological sciences including a strong foundation in physical sciences (chemistry, physics, calculus) and basic biological sciences. However, applicants with a diversity of other degrees also are successful. Courses in marine biology and oceanography are not necessary for entrance into the program. Applicants should contact specific faculty to discuss mutual research interests. The GRE score is not required for admission. Applicants whose first language is not English must pass the Test of English as a Foreign Language (TOEFL) with a score of at least 550. All application requirements are available  here .

Curriculum Requirements

Each student will design a customized curriculum based on their research focus with advisor approval.

Minimum of 18 course credits and 12 dissertation credits.

No core courses are required. Students choose courses with the advice of their committee. 

• At the end of the second year, a written qualifying examination is required of all Ph.D. students that focuses on the subject matter needed to complete the research proposed for the dissertation.
• The purpose of the qualifying examination is to demonstrate that the MBE doctoral student has the necessary understanding and expertise in research and related fields to complete the dissertation research. The topic areas should be agreed upon by the student, chair, and the dissertation committee soon after the proposal defense. The student is strongly encouraged to discuss the specific topics with each member of the dissertation committee, well in advance of the examination, to clarify the expected questions. The committee is encouraged to provide specific reading or areas of knowledge they will test the student on.
• The qualifying exam is 4 - 5 partial days (4 hours per day) on questions written by their committee. It is the advisor’s responsibility to provide the test and to have the student’s committee grade it in a timely manner. The scheduling of the exam sessions is the responsibility of the student's advisor, but in all cases the written portion of the examination shall be completed within one week.
• An additional oral qualifying examination may be required by the student's committee, but may not serve as a substitute for the written examination, which is a Graduate School requirement.
• The decision of passing or failing the qualifying examination rests with the dissertation committee. The qualifying examination (written and, if required, oral) must be successfully completed, as documented by the dissertation committee, before the student can be admitted to candidacy.
• In the event of a failure, a student may be re-examined once upon the recommendation of the student's committee in consultation with the academic committee. If permitted, the reexamination must be given before the end of the following semester.
• Attendance to the MBE seminars is required every semester. 
• Students entering the MBE Ph.D. program with a Master's degree begin to give seminars in their first year.
• Ph.D. students are expected to be a Teaching Assistant (TA) for two courses while pursuing their degree.
• The mandatory TA program will include training of new TAs, evaluation of their performance, and recognition of excellence. The goal is to make the experience as valuable as possible for the TA, the faculty, and the students taking our courses.
• A training session and two teaching opportunities are offered as courses in educational training (RSM 771, RSM 772, RSM 773). Students will be registered accordingly.
• Specific requirements for TAs are outlined in the Rosenstiel School Student Handbook.

Sample Plan of Study

The mission of the Marine Biology and Ecology (MBE) Ph.D. program is to train the next generation of scientists in the fundamental skills, knowledge, and practice of biology of the oceanic environment. Through coursework and independent research, we strive to prepare our students for positions in academia, government, or industry in jobs that leverage their skills in critical thinking, current technical knowledge such as statistical analysis and modeling, and understanding of the global marine environment. Our program commits to inspire graduates to continued scholarship, service, and innovation in an environment that is inclusive and diverse.

The goal is for Ph.D. students to demonstrate mastery of the fundamental skills, knowledge, and practice of biology of the oceanic environment, and commitment to scholarship, service, and innovation in an environment that is inclusive and diverse.

Student Learning Outcomes

• Students will demonstrate a broad understanding of marine and atmospheric science and an awareness of how scientific research in their topical areas bears on current human and societal issues.
• Students will be able to critically evaluate scientific literature, review previous knowledge on a topic, formulate testable hypotheses, and skillfully use available data and tools to advance knowledge in a topical area. They will be able to conduct high-quality, doctoral research as evidenced by their dissertation research.
• Students will demonstrate advanced oral and written communication skills, and be able to effectively communicate scientific information to a peer audience.

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Uniting the Marine Biological Laboratory and the University of Chicago research and education experience for an unparalleled program.

The newly established program between the Marine Biological Laboratory (MBL) and the University of Chicago (UChicago) leverages the unique partnership between two leading research institutions and combines the best of both worlds – access to a collaborative and expansive research environment that spans the scales of biological discovery at the MBL in Woods Hole, MA and the first-class resources of the University of Chicago. The formal affiliation and collaboration between these institutions was established in 2013 and has resulted in synergies in research and teaching that now expand to the graduate community.

Program Structure

Current MBL/UChicago graduate student opportunities are available through UChicago’s  Committee on Development, Regeneration, and Stem Cell Biology (DRSB) Program  and the Integrative Biology (IB) Program  with additional options available in the near future. Candidates should apply directly to the DRSB or IB Program depending on their specific interests. If accepted, students will complete their initial course requirements at UChicago before joining the MBL for summer research rotations in laboratories of affiliated faculty. Students will complete their thesis research at the MBL, with committee meetings, retreat participation and other activities encouraged through regular visits to Chicago throughout the graduate program.

Funding and Support

Students will be guaranteed full tuition, health insurance and generous stipend support. In addition, room and board will be provided during the first summer rotation at the MBL. After joining an MBL lab, students will be provided with funds to travel to Chicago for committee meetings and other events. Finally, students in the program will be eligible for an additional research allowance to be used at their discretion for research support or to attend meetings. Funds will also be available to students in UChicago labs to spend time at the MBL to foster collaborative research.

MBL Faculty Research Interests

MBL scientists are engaged in cutting-edge research   dedicated to scientific discovery in fundamental biology. There is a robust cadre of investigators with year-round laboratories, who are joined by hundreds of students and visiting researchers over the course of each year, making the Woods Hole community a unique environment to study, conduct research and engage with scientists from all over the world. Faculty affiliated with the DSRB and IB Programs conduct research across disciplines including cell biology, developmental and regenerative biology, neuroscience, sensory physiology, and comparative evolution and genomics. To learn more about the training faculty see the research interests of  Karen Echeverri ,  Andrew Gillis , Jennifer Morgan ,  Nipam Patel ,  Joshua Rosenthal , and Zak Swartz .

Karen Echeverri

Associate scientist, bell center.

The ability to regenerate complex tissue has fascinated scientists for a long time. The Echeverri lab is interested in the cell and molecular mechanisms driving regeneration. We study regeneration in axolotls; salamanders well known for their ability to functionally regenerate multiple body parts, including limbs, tail, heart, eyes and jaw and in addition can repair lesions in the brain and heal all wounds without forming scar tissue. My group uses cell and molecular tools combined with  in vivo  imaging and genomics approaches to decipher the key circuitry and cellular mechanisms that are essential to promote functional regeneration.

About Karen's work

Andrew Gillis

Associate scientist, bay paul center.

The Gillis Lab studies embryonic development of the skeletal, sensory, and neuroendocrine systems of vertebrate animals. Our work uses a comparative approach and a diverse array of emerging and established model organisms to reconstruct the origin and early evolution of the vertebrate body plan. Current lines of research include  1)  development, growth and repair of the vertebrate skeleton,  2)  serial homology and the origin of anatomical novelty, and  3)  the evolution of neuroendocrine cell type diversity.

Jennifer Morgan

Senior scientist, bell center.

The Morgan lab uses the sea lamprey ( Petromyzon marinus ) to elucidate mechanisms of nervous system degeneration and regeneration. Lampreys have a remarkable ability to regenerate axons and synapses after spinal injury, leading to functional recovery of swimming and other locomotor behaviors. We study these regenerative processes using transcriptomics, imaging, physiology and behavior. Moreover, lampreys possess a subset of giant synapses within the spinal cord that are amenable to acute molecular perturbations, allowing us to study synapse biology in health and disease. We are currently modeling how Parkinson’s and other neurodegenerative diseases impact synapses.

Nipam Patel

Mbl director.

My lab works to uncover both developmental and evolutionary mechanisms in a variety of animals that help us understand the generation of biodiversity, with a recent focus on the crustacean,  Parhyale , and several species of butterflies.  We use  Parhyale  to understand the role of Hox genes and other developmental genes in building and evolving body plans and also investigate the mechanisms that allows this species to regenerate its germline.  With butterflies, we are working to discover how single scale cells create the impressive nanostructures they use for structural coloration, without which they would be greatly restricted in the pallet of colors available to them.

Josh Rosenthal

Research in the Rosenthal Lab focuses on the editing of genetic information in mRNA. Rosenthal and collaborators discovered that cephalopods edit neuronal messages at unusually high levels, orders of magnitude more frequently than other organisms. Current research examines the mechanistic basis of high-level RNA editing in cephalopods, and how the process is influenced by environmental factors. In addition, the Rosenthal lab is leading efforts to create a genetically tractable cephalopod model at the MBL. In parallel to the cephalopod studies, the lab also focuses on the use of RNA editing as a platform for human therapeutics, partnering with biotech in Cambridge, MA and collaborative academic teams to edit away chronic pain and genetic mutations.

Assistant Scientist, Bell Center

Research in the Swartz Lab is focused on understanding how animals make eggs, and how eggs make animals. The laboratory is defining molecular mechanisms that underly the development of an egg from its precursor cell, the oocyte. To address these questions, they are working at the interface between cell biology and development, with the sea star Patiria miniata and its relatives. Sea stars produce millions of oocytes throughout their lives, and are amenable to an extensive range of molecular and genetic tools, as well as high resolution light microscopy. As relatives to the animal group that includes vertebrates like ourselves, the sea star can teach us important lessons relevant for human reproductive health, disease, and aging.

The PhD program in Marine and Environmental Sciences will train independent scientists whose research addresses fundamental and applied questions at the local, regional, national, and global scale.

Northeastern’s PhD program in Marine and Environmental Sciences trains high-caliber and independent scientists whose research addresses fundamental and applied marine and environmental science questions at local, regional, national, and global scales.  Our students will apply their research to address issues of relevance to society and the environment. In addition to training students to conduct world class research, we aim to prepare them to effectively communicate their findings broadly. We foster opportunities to develop teaching skills and communicate knowledge through outreach programs locally and internationally, so that students hone these essential skills.

To align their studies with their academic and career goals, students select one of the following 4 concentrations: Marine Sciences, Geosciences, Sustainability Sciences, and Ecology & Evolutionary Biology. While research interest may be focused, the academic experience is enriched by providing access to faculty with a wide range of expertise such as coastal sustainability, geology, marine biology, biogeochemistry, global change biology, marine robotics, fisheries science and policy, coral reef ecology, and social and ecological coupling.

• Bachelor’s and Advanced-degree entry are possible
• Students prepare and defend a written dissertation
• Faculty’s research areas are numerous
• State-of-the-art lab facilities located in Nahant, MA, a peninsula surrounded by the Atlantic Ocean
• Significant inter-lab collaboration encouraged and supported by the university
• Opportunities to TA for courses in Panamá and Washington state through the Three Seas Program

Our graduates pursue careers within academia and beyond.

• Tufts University
• Emory University
• University of Rhode Island
• Rutgers University
• Michigan State University
• New England Biolabs
• Knauss Fellowship Program

Application Materials

Application.

• Application fee – US $100
• Unofficial transcripts for all institutions attended (Official transcripts required upon acceptance of admission offer)
• Personal statement
• Three letters of recommendation
• GRE General – recommended, but not required
• Proof of English Proficiency for all applicants

Priority deadline for completed applications: December 1 st

Rolling admissions until March 15.  Check with department to see if there is availability.

• Program Website

Request Information for PhD in Marine and Environmental Sciences

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Marine Biology and Ecology

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• Graduate Programs

The Marine Biology and Ecology academic program focuses on a wide range of field, laboratory, and theoretical coursework in a range of research areas, such as coral reef studies, biological oceanography, and marine biology, fisheries biology, and the biology and behavior of marine animals.

MBE faculty, students, and their many international collaborators participate in multi-institutional, multi-disciplinary research programs. The UM Rosenstiel School hosts many research centers and groups, such as the experimental hatchery , The National Center for Coral Reef Research, the Rescue-a-Reef citizen science initiative , and the world-renowned NIH/University of Miami National Resource for  Aplysia .

Graduate students can choose from a diversity of research areas and coursework taught by internationally recognized scientists studying corals and climate change, fisheries biology, and conducting biomedical research.

Degrees Offered

• Doctor of Philosophy (Ph.D.)
• Master of Science (M.S.)
• Master of Professional Science (M.P.S.)   with tracks in Marine Mammal Science and Tropical Marine Ecosystem Management .

Requirements

Most successful applicants have a bachelor’s degree in biological sciences including a strong foundation in physical sciences (chemistry, physics, calculus) and basic biological sciences. However, applicants with a diversity of other degrees also are successful. Courses in marine biology and oceanography are not necessary for entrance into the program. Applicants should contact specific faculty to discuss mutual research interests. The GRE score is not required for admission. Applicants whose first language is not English must pass the Test of English as a Foreign Language (TOEFL) with a score of at least 550. All application requirements are available here .

Funding Opportunities

The majority of Ph.D. students are supported as Graduate Research Assistants during their tenure (typically 5 years). You can find more information on stipends, tuition, and health insurance   here . Students are also encouraged to apply for national fellowships. Additionally, students can apply or be nominated for the University of Miami and Rosenstiel School fellowships and awards .

Dr. Elizabeth Babcock Graduate Program Director [email protected]

Graduate Programs at the Rosenstiel School of Marine, Atmospheric, and Earth Science

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• University of Hawaiʻi at Mānoa

Marine Biology Graduate Program

Creating future leaders in marine science, welcome to the marine biology graduate program site, begin your adventure.

The Marine Biology Graduate Program is an interdisciplinary program co-administered by the College of Natural Sciences and the School of Ocean and Earth Sciences and Technology, offering MS and PhD degrees in Marine Biology. Marine biological studies at the University of Hawaiʻi have a long history of excellent research and graduate training in highly competitive programs like Botany, Microbiology, Oceanography, and Zoology. The Marine Biology Graduate Program provides graduate students access to experts in all of these disciplines. We encourage you to have a look and explore what we offer.

Megan Donahue and Celia Smith, Co-Directors

What we offer

The faculty and students in this program have access to one of largest coral reef habitats in the United States including the Main Hawaiian Islands, the Papahānaumokuākea Marine National Monument, and the U.S. Affiliated Pacific Islands. The unique location of the Hawaiian Archipelago in the central North Pacific Ocean makes it one of the most remote locations in the world allowing the processes that govern adaptation and evolution in the marine environment to be studied with minimal anthropogenic influence. The program emphasizes scientific training in marine biology as a high demand, interdisciplinary occupation for the future.

Degrees Available

• Master's Plan A in Marine Biology
• Doctorate in Marine Biology

Prospective graduate students may apply for either degree objective.

Undergraduate MB Degree

Although we are not affiliated with the undergraduate Marine Biology program, we provide a link to it here for your convenience.

Sponsorship

Identification of a sponsor is not required to apply. However, students must be sponsored by a Marine Biology faculty member to be admitted. Thus, it is in the applicant's best interest to contact potential sponsors well before the application deadline to discuss research intents & goals and to ask if they can provide research space and mentoring during your graduate study.

Prospective Students

Students are expected to have strong undergraduate or MS training in biological oceanography, biology, botany, environmental science, marine biology, microbiology, zoology, or other life sciences. We encourage you to prepare academically before applying to the program, as the time required to take courses in order to have the necessary knowledge after enrollment may hinder your graduate progress in the program.

Our partners

This program takes advantage of the 40+ faculty members whose research interests span the study of marine ecosystems, marine biogeochemical processes, reef and oceanic fisheries, and human/marine interactions. We also partner with state and federal agencies such as the Pacific Islands Fisheries Science Center for NOAA, the Hawaiʻi Department of Land and Natural Resources, and the Department of Business, Economic Development, and Tourism to offer research and internship opportunities.

Discover more

We invite you to explore our site and learn more about what the Program has to offer.

• Admissions Requirements Review all requirements for getting into the Program.
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• News and Events Go here for up to date info on current events, job and volunteer opportunities, and announcements.
• Contact Us Feel free to reach out to us for additional information.

"Don't wait any longer. Dive in the ocean, Leave and let the sea be you." Rumi

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Graduate Study in Marine Biology

Nearly three quarters of the Earth is covered by an ocean that harbors most of life’s diversity, feeds and powers coastal economies, regulates global climate, and is a rich source of biomedical models, natural products, and new drugs. The Marine Biology program offers excellent opportunities to pursue graduate research in areas such as the evolution, development, and sensory biology of marine organisms, microbiology, ichthyology, and marine conservation science. Prospective students are encouraged to explore the research interests of our faculty and then directly contact those professors with whom they might like to work, in addition to submitting an application to the graduate school.

Marine Biology students begin their curriculum with a fall semester consisting of four back-to-back, one-monthlong intensive research courses.  The graduate curriculum for Marine Biology students is highly flexible and is customized to the needs and interests of the individual student.  By the end of their second year, PhD students complete a written preliminary examination that tests general knowledge in marine biology, and an oral qualifying exam that focuses on areas critical to the student’s dissertation research.

Marine Biology graduate students benefit from a strongly interdisciplinary marine science program , involving the Departments of Biology and  Earth & Environment , as well as close partnerships with the New England Aquarium , National Marine Sanctuary Program , and Sea Education Association .

Check the list of recent dissertations and publications to see the range of topics addressed by Marine Biology students.

Interested in applying to our graduate program? 

All applicants should review the application procedures listed on our apply page . MS applicants please note that we anticipate limited space available in the Fall 2024 Marine Semester; those applying for the MS with coursework or scholarly paper tracks in Marine Biology should plan alternate fall courses or delay application to a future year. 

MB applicants interested in applying to our MS (Coursework Track or Scholarly Paper Track) do not need to contact faculty before applying. 

MB applicants interested in applying for our PhD or MS (Research Track) require that you have a professor who is willing to accept you into his/her/their lab. We strongly recommend that you contact professors with whom you are interested in working before you apply to the MB graduate program. Below are guidelines on how to contact professors via e-mail. 

• Locate current research projects and contact information for professors in MB who are currently accepting students (see below).
• Your research interests and how they align with the research of the faculty member (1-2 sentences).
• Your previous research experience (2-3 sentences). 
• A new idea you might like to pursue while working with the lab you are contacting. This research idea should be consistent with the lab’s current research capabilities and demonstrate some degree of familiarity with the field (2-3 sentences).
• Responses should be as text in e-mail, not as an attachment. However, please attach your resume/curriculum vitae to your e-mail.

List of MB faculty currently accepting PhD students for Fall 2024:

Peter Buston , Associate Professor of Biology; Director of the BU Marine Program

Evolutionary Ecology, Animal Behavior, Marine Ecology and Biological Oceanography.

Ethan Deyle , Research Assistant Professor of Biology

Quantitative Ecology, Environmental Data Science, Nonlinear Dynamics, Applied Complex Systems, Marine Ecology

Robinson W. Fulweiler , Professor of Biology (jointly with Earth & Environment)

Biogeochemistry and Marine Ecology

List of MB faculty currently accepting MS students for Fall 2024:

John R. Finnerty , Associate Professor of Biology

Evolutionary and ecological developmental biology; evolutionary and ecological genomics; marine biodiversity; global change biology; coral conservation.

Phillip S. Lobel , Professor of Biology

ichthyology; behavioral ecology and taxonomy of fishes

Randi Rotjan , Research Associate Professor of Biology; Senior Lecturer; Director of Masters Studies

Marine ecology, conservation biology, behavioral ecology, organismal physiology, coral reefs

Visit our MB Research page for a list of all of the EBE faculty.

Integrative, Comparative & Marine Biology, Ph.D.

Among Public National Universities, U.S. News & World Report

Featured in the Fiske Guide to Colleges

Nationally, Best Online Bachelor's Programs, U.S. News & World Report

Our alumni work in:

Conduct research at a university or college.

Use your knowledge and skills to conduct research for government organizations.

Design and develop technologies to research and solve marine-related issues.

Influence marine policy with your advanced knowledge and skills.

Become faculty at a university or college.

Develop a research plan

Present and defend a research plan

Answer questions regarding their research field

Communicate his or her research to a broadly trained public audience

Write research in the form of a manuscript for publication in a scientific journal

“ The department has a wide array of interests, especially centered around the marine environment, that cultivate a rich academic community. With the addition of very friendly faculty and students, it's a hard place to leave. ”

Advance Your Research

The Doctor of Philosophy program offers a broad background and overview of the fields comprising marine biology and prepares you for professional and leadership positions.

This research degree program serves students with interests in conducting research in academia, industry and government; individuals who intend to become faculty in undergraduate teaching institutions; managers in technology-based industries and policy makers in government.

Interested in teaching? The program includes a teaching practicum with classroom instruction in pedagogical techniques and technologies along with lecture experience under the guidance of a faculty mentor.

Diverse Research Areas

Choose from diverse research areas and coursework that provide a broad educational perspective including coastal and estuarine biology, crustacean biology, marine mammalogy, and molecular biology and systematics of marine organisms.

Sample Courses in Integrative, Comparative & Marine Biology, Ph.D.

How do i apply to the integrative, comparative & marine biology, ph.d. program.

You are required to submit the following to the Graduate School (please do not send items to the department):

• An application for graduate admission
• Official transcripts of all college work (undergraduate and graduate)
• Three recommendations by individuals in professionally relevant fields
• Current curriculum vitae
• Detailed summary of M.S. thesis research (maximum of three pages)
• Statement of interest for Ph.D. research (maximum of three pages)
• Reprints or copies of any publications (if applicable)

Admission Deadlines & Requirements

Information: ph.d. integrative, comparative, and marine biology.

• Complete applications are reviewed for admission by the program as they are received.
• All application and supporting documents must be received by the published deadline.

Coordinator

• Dr. Stephanie Kamel 910-962-2841 [email protected]  

Location/Delivery Method

• Main Campus

Deadlines (11:59 p.m. EST)

• Deadline: October 15, 2023
• Deadline:  April 15, 2024
• Priority Consideration: February 15, 2024
• Space Available Deadline: May 15, 2024

Transcripts

• One official transcript is required from each U.S. post-secondary institution attended. Refer to the Getting Started page for international transcript instructions.

Test Scores

• None Required

Recommendations

• 3 recommendation letters by individuals in professionally relevant fields.

Additional Requirements

• Upload Supplemental Documents After Application Submission Upload Supplemental Documents
• Curriculum Vitae
• Ph.D. Research:   Statement of Interest for Ph.D. research, including a summary of relevant experience (maximum of three pages)
• Publications:  Copies of publications (if available)
• Degree Requirements:   A bachelor’s degree in biology or a related discipline from a regionally accredited U.S. college or university or its equivalent at an international institution based on a four-year program.
• Research Requirements:   A record of research, applied learning or relevant professional experience.
• UNCW Faculty: Prospective students should make initial contact with the faculty with whom you wish to work  (Meet Our Team - Faculty).
• Note:   Under certain circumstances, a student in the M.S. biology or marine biology program may, with the support of their faculty advisor, apply to the Ph.D. program rather than completing their M.S. degree. Students who choose this path after their first year of core courses and research planning must complete a new application, including letters of recommendation, to enter the Ph.D. program.

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Related Programs

Coastal and ocean policy, m.s..

The Master of Coastal and Ocean Policy (MCOP) is a professional, non-thesis, interdisciplinary degree program that immerses students in a unique curriculum aimed at advancing technical knowledge of coastal and ocean processes and resources as well as the tools and concepts of the policy making.

Geographic Information Science, Post-Baccalaureate Certificate

UNCW is accredited with the United States Geospatial Intelligence Foundation (USGIF) to offer a certificate in Geospatial Intelligence (GEOINT).

Graduate Programs

Graduate programs to consider, biology, m.s..

Specialize in ocean sciences, technology and marine policy.

Marine Biology, M.S.

A program that prepares students as managers of coastal and marine resources, trained to deal with contemporary problems in the environment; and provides a broad–based graduate program allowing for specialization in the diverse fields of inquiry under the marine biology umbrella.

Academic Catalog 2023-2024

Marine and environmental sciences, phd.

The PhD in Marine and Environmental Sciences (MES) program provides students with advanced course work and training in the concentration areas of marine sciences, geosciences, sustainability sciences, and ecology and evolutionary biology. 

Students must pass three examinations during the course of their graduate studies:

• An oral examination by the student’s dissertation committee.
• A proposal defense presented to the student's dissertation committee that explains the research areas that the student proposes to work in.
• A defense of the student's written dissertation consisting of a public seminar, public question-and-answer period, and private defense of their work to their dissertation committee. Dissertation committees consist of at least four Northeastern faculty and one external faculty member.

A cumulative GPA of 3.000 is required for graduation. All PhD students are required to have at least two first-authored publications submitted to or accepted in a peer-reviewed journal prior to their defense. The PhD will be awarded following submission of a dissertation, approved by the candidate’s dissertation committee, to the College of Science.

Students who ​do not qualify for the doctoral degree, but who have completed ​required coursework with a cumulative GPA of 3.000 or better, may be eligible to receive ​a terminal  MS Marine and Environmental Sciences  degree. Note that no students will be admitted directly into the Marine and Environmental Sciences program to pursue a masterʼs degree.

Bachelor's Degree Entrance

Complete all courses and requirements listed below unless otherwise indicated.

Annual review Dissertation committee Qualifying examination Dissertation proposal Candidacy First-author publication Dissertation defense

Core Requirements

Concentration.

Complete one of the following concentrations:

Ecology and Evolutionary Biology

Sustainability sciences, geosciences, marine sciences, dissertation, program credit/gpa requirements.

30 total semester hours required Minimum 3.000 GPA required

10 total semester hours required Minimum 3.000 GPA required

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Texas A&M University Catalogs

Doctor of philosophy in marine biology.

The PhD degree in Marine Biology focuses on independently supervised research complemented by formal coursework. Essential components of the program include a diverse curriculum focusing on ecology and evolution of marine organisms and quantitative analysis; and original, supervised scholarly research in marine biology, to be written and formally defended as a dissertation.

Steps to Fulfill a Doctoral Program

Program Requirements

• Student's Advisory Committee

Degree Plan

Transfer of credit, research proposal.

• Preliminary Examination

Preliminary Examination Format

Preliminary examination scheduling, report of preliminary examination, failure of the preliminary examination, retake of failed preliminary examination.

• Final Examination

Report of Final Examination

Dissertation, student’s advisory committee.

After receiving admission to graduate studies and enrolling, the student will consult with the head of his or her major or administrative department (or chair of the intercollegiate faculty) concerning appointment of the chair of the advisory committee. The student’s advisory committee will consist of  no fewer than four members of the graduate faculty  representative of the student’s several fields of study and research, where the chair or co-chair must be from the student’s department (or intercollegiate faculty, if applicable), and  at least one or more of the members must have an appointment to a department other than the student’s major department . The outside member for a student in an interdisciplinary degree program must be from a department different from the chair of the student’s committee.

The chair, in consultation with the student, will select the remainder of the advisory committee. Only graduate faculty members located on Texas A&M University campuses may serve as chair of a student’s advisory committee. Other Texas A&M University graduate faculty members located off-campus may serve as a member or co-chair (but not chair), with a member as the chair.

If the chair of a student’s advisory committee voluntarily leaves the University and the student is near completion of the degree and wants the chair to continue to serve in this role, the student is responsible for securing a current member of the University Graduate Faculty, from the student’s academic program and located near the Texas A&M University campus site, to serve as the co-chair of the committee. The Department Head or Chair of Intercollegiate faculty may request in writing to the Associate Provost and Dean of the Graduate and Professional School that a faculty member who is on an approved leave of absence or has voluntarily separated from the university, be allowed to continue to serve in the role of chair of a student’s advisory committee without a co-chair for up to one year. The students should be near completion of the degree. Extensions beyond the one year period can be granted with additional approval of the Dean.

The committee members’ signatures on the degree plan indicate their willingness to accept the responsibility for guiding and directing the entire academic program of the student and for initiating all academic actions concerning the student. Although individual committee members may be replaced by petition for valid reasons, a committee cannot resign  en masse . The chair of the committee, who usually has immediate supervision of the student’s research and dissertation or record of study, has the responsibility for calling all meetings of the committee. The duties of the committee include responsibility for the proposed degree plan, the research proposal, the preliminary examination, the dissertation or record of study and the final examination. In addition, the committee, as a group and as individual members, is responsible for counseling the student on academic matters, and, in the case of academic deficiency, initiating recommendations to the Graduate and Professional School.

The student’s advisory committee will evaluate the student’s previous education and degree objectives. The committee, in consultation with the student, will develop a proposed degree plan and outline a research problem which, when completed, as indicated by the dissertation (or its equivalent for the degree of Doctor of Education or the degree of Doctor of Engineering), will constitute the basic requirements for the degree. The degree plan must be filed with the Graduate and Professional School prior to the deadline imposed by the student’s college and no later than 90 days prior to the preliminary examination.

This proposed degree plan should be submitted through the online Document Processing Submission System located on the website  http://ogsdpss.tamu.edu . A minimum of 60 hours is required on the degree plan for the Doctor of Philosophy for a student who has completed a master’s degree. A student who has completed a DDS/DMD, DVM or a MD at a U.S. institution is also required to complete a minimum of 60 hours. A student who has completed a baccalaureate degree but not a master’s degree will be required to complete a 90-hour degree plan. Completion of a DDS/DMD, DVM or MD degree at a foreign institution requires completion of a minimum of 90 hours for the Doctor of Philosophy. A field of study may be primarily in one department or in a combination of departments. A degree plan must carry a reasonable amount of 691 (research). A maximum of 9 hours of 400-level undergraduate courses may be used toward meeting credit-hour requirements for the Doctor of Philosophy.

Additional coursework may be added by petition to the approved degree plan by the student’s advisory committee if it is deemed necessary to correct deficiencies in the student’s academic preparation. No changes can be made to the degree plan once the student’s Request for Final Examination is approved by the Graduate and Professional School.

Approval to enroll in any professional course (900-level) should be obtained from the head of the department (or Chair of the intercollegiate faculty, if applicable) in which the course will be offered before including such a course on a degree plan.

No credit may be obtained by correspondence study, by extension or for any course of fewer than three weeks duration.

For non-distance degree programs, no more than four courses may be taken by distance education without approval of the Graduate and Professional School and no more than 50 percent of the non-research credit hours required for the program may be completed through distance education courses.

To receive a graduate degree from Texas A&M University, students must earn one-third or more of the credits through the institution’s own direct instruction. This limitation also applies to joint degree programs. 

Courses for which transfer credits are sought must have been completed with a grade of B or greater and must be approved by the student’s advisory committee and the Graduate and Professional School. These courses must not have been used previously for another degree. Except for officially approved cooperative doctoral programs, credit for thesis or dissertation research or the equivalent is not transferable. Credit for “internship” coursework in any form is not transferable. Courses taken in residence at an accredited U.S. institution or approved international institution with a final grade of B or greater will be considered for transfer credit if, at the time the courses were completed, the courses would be accepted for credit toward a similar degree for a student in degree-seeking status at the host institution. Credit for coursework taken by extension is not transferable. Coursework  in which no formal grades are given or in which grades other than letter grades (A or B) are earned (for example, CR, P, S, U, H, etc.) is not accepted for transfer credit . Credit for coursework submitted for transfer from any college or university must be shown in semester credit hours, or equated to semester credit hours.

Courses used toward a degree at another institution may not be applied for graduate credit. If the course to be transferred was taken prior to the conferral of a degree at the transfer institution, a letter from the registrar at that institution stating that the course was not applied for credit toward the degree must be submitted to the Graduate and Professional School.

Grades for courses completed at other institutions are not included in computing the GPA. An official transcript from the university at which transfer courses are taken must be sent directly to the Office of Admissions.

The general field of research to be used for the dissertation should be agreed on by the student and the advisory committee at their first meeting, as a basis for selecting the proper courses to support the proposed research.

As soon thereafter as the research project can be outlined in reasonable detail, the dissertation research proposal should be completed. The research proposal should be approved at a meeting of the student’s advisory committee, at which time the feasibility of the proposed research and the adequacy of available facilities should be reviewed. The approved proposal, signed by all members of the student’s advisory committee, the head of the student’s major department (or chair of the intercollegiate faculty, if applicable), must be submitted to the Graduate and Professional School at least 20 working days prior to the submission of the Request for the Final Examination.

Compliance issues must be addressed if a graduate student is performing research involving human subjects, animals, infectious biohazards and recombinant DNA. A student involved in these types of research should check with the Office of Research Compliance and Biosafety at (979) 458-1467 to address questions about all research compliance responsibilities. Additional information can also be obtained on the website  http:// rcb.tamu.edu .

Examinations

Preliminary examination for doctoral students.

The student’s major department (or chair of the interdisciplinary degree program faculty, if applicable) and his or her advisory committee may require qualifying, cumulative or other types of examinations at any time deemed desirable. These examinations are entirely at the discretion of the department and the student’s advisory committee.

The preliminary examination is required. The preliminary examination for a doctoral student shall be given no earlier than a date at which the student is within 6 credit hours of completion of the formal coursework on the degree plan (i.e., all coursework on the degree plan except 681, 684, 690, 691, 692, 693, 695, 697, 791, or other graduate courses specifically designated as S/U in the course catalog). The student should complete the Preliminary Examination no later than the end of the semester following the completion of the formal coursework on the degree plan.

The objective of preliminary examination is to evaluate whether the student has demonstrated the following qualifications:

a.     a mastery of the subject matter of all fields in the program;

b.     an adequate knowledge of the literature in these fields and an ability to carry out bibliographical research;

c.     an understanding of the research problem and the appropriate methodological approaches.

The format of the preliminary examination shall be determined by the student’s department (or interdisciplinary degree program, if applicable) and advisory committee, and communicated to the student in advance of the examination. The exam may consist of a written component, oral component, or combination of written and oral components.

The preliminary exam may be administered by the advisory committee or a departmental committee; herein referred to as the examination committee.

Regardless of exam format, a student will receive an overall preliminary exam result of pass or fail. The department (or interdisciplinary degree program, if applicable) will determine how the overall pass or fail result is determined based on the exam structure and internal department procedures. If the exam is administered by the advisory committee, each advisory committee member will provide a pass or fail evaluation decision.

Only one advisory committee substitution is allowed to provide an evaluation decision for a student’s preliminary exam, and it cannot be the committee chair.

If a student is required to take, as a part of the preliminary examination, a written component administered by a department or interdisciplinary degree program, the department or interdisciplinary degree program faculty must:

a.     offer the examination at least once every six months. The departmental or interdisciplinary degree program examination should be announced at least 30 days prior to the scheduled examination date.

b.     assume the responsibility for marking the examination satisfactory or unsatisfactory, or otherwise graded, and in the case of unsatisfactory, stating specifically the reasons for such a mark.

c.     forward the marked examination to the chair of the student’s advisory committee within one week after the examination.

Prior to commencing any component of the preliminary examination, a departmental representative or the advisory committee chair will review the eligibility criteria with the student, using the Preliminary Examination Checklist to ensure the student is eligible for the preliminary examination. The following list of eligibility requirements applies.

Student is registered at Texas A&M University for a minimum of one semester credit hour in the long semester or summer term during which any component of the preliminary examination is held. If the entire examination is held between semesters, then the student must be registered for the term immediately preceding the examination.

An approved degree plan is on file with the Graduate and Professional School prior to commencing the first component of the examination.

Student’s cumulative GPA is at least 3.000.

Student’s degree plan GPA is at least 3.000.

At the end of the semester in which at least the first component of the exam is given, there are no more than 6 hours of coursework remaining on the degree plan (except 681, 684, 690, 691, 692, 693, 695, 697, 791, or other graduate courses specifically designated as S/U in the course catalog). The head of the student’s department (or Chair of the Interdisciplinary Degree Program, if applicable) has the authority to approve a waiver of this criterion.

Credit for the preliminary examination is not transferable in cases where a student changes degree programs after passing a preliminary exam.

If a written component precedes an oral component of the preliminary exam, the chair of the student’s examination committee is responsible for making all written examinations available to all members of the committee.  A positive evaluation of the preliminary exam by all members of a student’s examination committee with at most one dissension is required to pass a student on his or her preliminary exam.

The student’s department will promptly report the results of the Preliminary Examination to the Graduate and Professional School via the Report of Doctoral Preliminary Examination form. The Preliminary Examination checklist form must also be submitted. These forms should be submitted to the Graduate and Professional School within 10 working days of completion of the preliminary examination.

The Report of the Preliminary Examination form must be submitted with original signatures of the approved examination committee members. If an approved examination committee member substitution (one only) has been made, that signature must also be included, in place of the committee member, on the form submitted to the Graduate and Professional School. The original signature of the department head is also required on the form.

After passing the required preliminary oral and written examinations for a doctoral degree, the student must complete the final examination within four years of the semester in which the preliminary exam is taken. Exams taken in between terms will expire at the end of the term that ended prior to the exam. For example, a preliminary exam taken and passed during the fall 2019 semester will expire at the end of the fall 2023 semester. A preliminary exam taken in the time between the summer and fall 2019 semesters will expire at the end of the summer 2023 semester.

First Failure

Upon approval of a student’s examination committee (with no more than one member dissenting), and approval of the Department and Graduate and Professional School, a student who has failed a preliminary examination may be given one re-examination. In accordance with Student Rule 12.5, the student’s department head or designee, intercollegiate faculty, or graduate advisory committee should make a recommendation to the student regarding their scholastic deficiency.

Second Failure

Upon failing the preliminary exam twice in a doctoral program, a student is no longer eligible to continue to pursue the PhD in that program/major. In accordance with Student Rule 12.5.3 and/or 12.5.4, the student will be notified of the action being taken by the department as a result of the second failure of the preliminary examination.

Adequate time must be given to permit a student to address inadequacies emerging from the first preliminary examination. The examination committee must agree upon and communicate to the student, in writing, an adequate time-frame from the first examination (normally six months) to retest, as well as a detailed explanation of the inadequacies emerging from the examination. The student and committee should jointly negotiate a mutually acceptable date for this retest.  When providing feedback on inadequacies, the committee should clearly document expected improvements that the student must be able to exhibit in order to retake the exam.  The examination committee will document and communicate the time-frame and feedback within 10 working days of the exam that was not passed.

Final Examination for Doctoral Students

The candidate for the doctoral degree must pass a final examination by deadline dates announced in the Graduate and Professional School Calendar each semester. The doctoral student is allowed only one opportunity to take the final examination.

No unabsolved grades of D, F, or U for any course can be listed on the degree plan. The student must be registered for any remaining hours of 681, 684, 690, 691, 692, 791 or other graduate courses specifically designated as S/U in the course catalog during the semester of the final exam. No student may be given a final examination until they have been admitted to candidacy and their current official cumulative and degree plan GPAs are 3.00 or better.

To be admitted to candidacy for a doctoral degree, a student must have:

1.       completed all formal coursework on the degree plan with the exception of any remaining 681, 684, 690 and 691, 692 (Professional Study), or 791 hours,

2.       a 3.0 Graduate GPA and a Degree Plan GPA of at least 3.0 with no grade lower than C in any course on the degree plan,

3.       passed the preliminary examination,

4.       submitted an approved dissertation proposal,

5.       met the residence requirements.

The request to hold and announce the final examination must be submitted to the Graduate and Professional School a minimum of 10 working days in advance of the scheduled date. Any changes to the degree plan must be approved by the Graduate and Professional School prior to the submission of the request for final examination.

 The student’s advisory committee will conduct this examination. The final examination is not to be administered until the dissertation or record of study is available in substantially final form to the student’s advisory committee, and all concerned have had adequate time to review the document.  Whereas the final examination may cover the broad field of the candidate’s training, it is presumed that the major portion of the time will be devoted to the dissertation and closely allied topics. Persons other than members of the graduate faculty may, with mutual consent of the candidate and the chair of the advisory committee, be invited to attend a final examination for an advanced degree. A positive vote by all members of the graduate committee with at most one dissension is required to pass a student on his or her exam. A department can have a stricter requirement provided there is consistency within all degree programs within a department. Upon completion of the questioning of the candidate, all visitors must excuse themselves from the proceedings.

The student’s department will promptly report the results of the Final Examination to the Graduate and Professional School via the Report of Doctoral Final Examination form. These forms should be submitted to the Graduate and Professional School within 10 working days of completion of the final examination. The Graduate and Professional School must be notified in writing of any cancellations.

A positive evaluation of the final exam by all members of a student’s advisory committee with at most one dissension is required to pass a student on his or her final exam. The Report of the Final Examination Form must be submitted with original signatures of only the committee members approved by the Graduate and Professional School. If necessary, multiple copies of the form may be submitted with different committee member original signatures. If an approved committee member substitution (1 only) has been made, his/her signature must be included on the form submitted to the Graduate and Professional School.

The ability to perform independent research must be demonstrated by the dissertation,  which must be the original work of the candidate . Whereas acceptance of the dissertation is based primarily on its scholarly merit, it must also exhibit creditable literary workmanship. The format of the dissertation must be acceptable to the Graduate and Professional School. Guidelines for the preparation of the dissertation are available in the  Thesis Manual , which is available online at  http://grad.tamu.edu .

After successful defense and approval by the student’s advisory committee and the head of the student’s major department (or chair of the intercollegiate faculty, if applicable), a student must submit his/her dissertation in electronic format as a single PDF file. The PDF file must be uploaded to the website,  http://grad.tamu.edu . Additionally, a signed paper approval form with original signatures must be received by the Graduate and Professional School. Both the PDF file and the signed approval form are required by the deadline.

Deadline dates for submitting are announced each semester or summer term in the Graduate and Professional School Calendar (see Time Limit statement). These dates also can be accessed via the website  http://grad.tamu.edu .

Each student who submits a document for review is assessed a one-time thesis/dissertation processing fee through Student Business Services. This processing fee is for the thesis/dissertation services provided. After commencement, dissertations are digitally stored and made available through the Texas A&M Libraries.

A dissertation that is deemed unacceptable by the Graduate and Professional School because of excessive corrections will be returned to the student’s department head or chair of the intercollegiate faculty . The manuscript must be resubmitted as a new document, and the entire review process must begin anew. All original submittal deadlines must be met during the resubmittal process in order to graduate.

Additional Requirements

Continuous registration, admission to candidacy.

• 99-Hour Cap on Doctoral Degree

Application for Degree

A student who enters the doctoral degree program with a baccalaureate degree must spend one academic year plus one semester in resident study at Texas A&M University. A student who holds master’s degree when he/she enters doctoral degree program must spend one academic year in resident study. One academic year may include two adjacent regular semesters or one regular semester and one adjacent 10-week summer semester. The third semester is not required to be adjacent to the one year. Enrollment for each semester must be a minimum of 9 credit hours each to satisfy the residence requirement. A minimum of 1 credit hour must be in a non-distance education delivery mode. Semesters in which the student is enrolled in all distance education coursework will not count toward fulfillment of the residence requirement.

To satisfy the residence requirement, the student must complete a minimum of 9 credit hours per semester or 10-week summer semester in resident study at Texas A&M University for the required period. A student who enters a doctoral degree program with a baccalaureate degree may fulfill residence requirements in excess of one academic year (18 credit hours) by registration during summer sessions or by completion of a less-than-full course load (in this context a full course load is considered 9 credit hours per semester).

Students who are employed full-time while completing their degree may fulfill total residence requirements by completion of less-than-full time course loads each semester. In order to be considered for this, the student is required to submit a Petition for Waivers and Exceptions along with verification of his/her employment to the Graduate and Professional School. An employee should submit verification of his/her employment at the time he/she submits the degree plan. See  Registration.

See  Residence Requirements .

All requirements for doctoral degrees must be completed within a period of ten consecutive calendar years for the degree to be granted. A course will be considered valid until 10 years after the end of the semester in which it is taken. Graduate credit for coursework more than ten calendar years old at the time of the final oral examination may not be used to satisfy degree requirements.

A final corrected version of the dissertation or record of study in electronic format as a single PDF file must be cleared by the Graduate and Professional School within one year of the semester in which the final exam is taken. Exams taken in between terms will expire at the end of the term that ended prior to the exam. For example, a final exam taken and passed during the fall 2022 semester will expire at the end of the fall 2023 semester. A final exam taken in the time between the summer and fall 2022 semesters will expire at the end of the summer 2023 semester. Failure to do so will result in the degree not being awarded.

A student in a program leading to a Doctor of Philosophy who has completed all coursework on his/her degree plan other than 691 (research) are required to be in continuous registration until all requirements for the degree have been completed. See  Continuous Registration Requirements .

• completed all formal coursework on the degree plan with the exception of any remaining 681, 684, 690 and 691 or 791.
• a 3.0 Graduate GPA and a Degree Plan GPA of at least 3.0 with no grade lower than C in any course on the degree plan,
• passed the preliminary examination (written and oral portions),
• submitted an approved dissertation proposal,
• met the residence requirements. The final examination will not be authorized for any doctoral student who has not been admitted to candidacy.

A student is required to possess a competent command of English. For English language proficiency requirements, see the Admissions section of this catalog. The doctoral (PhD) foreign language requirement at Texas A&M University is a departmental option, to be administered and monitored by the individual departments of academic instruction.

99-Hour Cap on Doctoral Degrees

In Texas, public colleges and universities are funded by the state according to the number of students enrolled. In accordance with legislation passed by the Texas Legislature, the number of hours for which state universities may receive subvention funding at the doctoral rate for any individual is limited to 99 hours. Texas A&M and other universities will not receive subvention for hours in excess of the limit.

Institutions of higher education are allowed to charge the equivalent of non-resident tuition to a resident doctoral student who has enrolled in 100 or more semester credit hours of doctoral coursework.

Doctoral students at Texas A&M have seven years to complete their degree before being charged out-of-state tuition. A doctoral student who, after seven years of study, has accumulated 100 or more doctoral hours will be charged tuition at a rate equivalent to out-of-state tuition. Please note that the tuition increases will apply to Texas residents as well as students from other states and countries who are currently charged tuition at the resident rate. This includes those doctoral students who hold GAT, GANT, and GAR appointments or recipients of competitive fellowships who receive more than $1,000 per semester. Doctoral students who have not accumulated 100 hours after seven years of study are eligible to pay in-state tuition if otherwise eligible.

Doctoral students who exceed the credit limit will receive notification from the Graduate and Professional School during the semester in which they are enrolled and exceeding the limit in their current degree program. The notification will explain that the State of Texas does not provide funding for any additional hours in which a student is enrolled in excess of 99 hours. Texas A&M University will recover the lost funds by requiring students in excess of 99 hours to pay tuition at the non-funded, non-resident rate. This non-funded, non-resident tuition rate status will be updated for the following semester and in all subsequent semesters until receipt of a doctoral degree. Please see the  Tuition Calculator  at the non-resident rate for an example of potential charges.

The following majors are exempt from the 99-Hour Cap on Doctoral Degrees and have a limit of 130 doctoral hours:

• Biochemistry
• Biomedical Sciences
• Clinical Psychology
• Counseling Psychology
• Genetics and Genomics
• Health Services Research
• Medical Sciences
• Microbiology
• Neurosciences (School of Medicine)
• Oral and Craniofacial Biomedical Sciences
• Pharmaceutical Sciences
• Public Health Sciences
• School Psychology

For information on applying for your degree, please visit the  Graduation  section.

• Apply to UMaine

School of Marine Sciences

Marine biology.

The M.S. and Ph.D. in Marine Biology are designed to provide students with a broad knowledge base in marine biology in areas including, but not restricted to, taxonomy/systematics, comparative morphology, evolution, genetics, physiology, cellular and molecular biology, and ecology. Marine Biology is a dynamic field, and our programs are flexibly designed to accommodate new discoveries and technologies.

Course requirements for all students are SMS 500 (Marine Biology), one semester of statistics at the graduate level, one semester of seminar (SMS 691) and a basic knowledge of descriptive oceanography.  Students interact closely with their committees to select additional coursework that provides both a broad knowledge base in marine biology and specific emphases appropriate for their research (e.g., aquaculture, biological oceanography, biology of fishes, ecology, invertebrate biology, marine algae, molecular biology). For more degree requirements, consult the Handbook for Graduate Programs  and the Marine Biology Milestone Tracking form .

Required Core Courses

To maintain maximum flexibility, the majority of courses comprising a student’s Program of Study will be determined by the student in consultation with his/her major advisor and advisory committee. Requirements include:

• Basic knowledge of descriptive oceanography equivalent to SMS 302.  Coursework taken prior to matriculation needs to be approved by the Graduate Program Coordinator.
• SMS 500 – Marine Biology Credits: 4
• SMS 691 – Marine Science Seminar Credits: 1
• Participation in SMS Graduate Student Symposium during each year of full-time study.
• One semester of graduate level statistics.

Elective Marine Biology Courses

• INT 510 – (BSC, SMS) Marine Invertebrate Zoology Credits: 5
• INT 563 – (BSC, SMS) Marine Benthic Ecology Credits: 3
• SMS 501 – Biological Oceanography Credits: 3
• SMS 514 – Ecology of Marine Sediments Credits: 2-3
• SMS 525 – Marine Biogeochemistry Credits: 3
• SMS 531 – Coral Reefs Credits: 3
• SMS 550 – Fisheries Oceanography Credits: 3
• SMS 562 – Fisheries Population Dynamics Credits: 3
• SMS 598 – Special Topics in Marine Science Credits: 1-3

Program of Study

Students will be expected to develop special expertise in an area of marine biology.  A Program of Study with additional coursework will be established in close consultation with the student’s advisor and committee to build the foundation of the student’s expertise.

Program Coordinator

Damian Brady Assistant Professor 193 Clarks Cove Road Walpole, ME  04573 207.581.5117 [email protected]

EEMB Doctor of Philosophy Degree

Ucsb course catalog.

View full list of 2023-24 EEMB courses.

PhD Graduate Handbook

Phd graduate checklist, foundation building.

To ensure that students have the fundamentals of the disciplines of Ecology, Evolution, and Marine Biology, students are required to take EEMB 508 and EEMB 509 in the fall and winter of their first year. UCSB is a highly interdisciplinary campus, and students are encouraged to take advantage of graduate level courses and seminars in EEMB and in other departments to further develop their knowledge of their own subject area and others of interest. Being a teaching assistant helps cement a doctoral student's knowledge and develops essential teaching and communication skills. Course credit is awarded for TAing and one course is required in teaching techniques.

Course Requirements

These seminar courses aredesigned to familiarize new graduate students with the EEMB faculty and the diversity of research being conducted in the department and to provide a forum for interactions among new graduate students. It is required of all incoming graduate students unless special circumstances prevent attendance.

All students must qualify for and hold a teaching assistantship for the equivalent of two quarters during some point in the graduate career in order to obtain teaching experience. To qualify, students must complete the following courses.

Graduate Council Regulations Regarding Committees

• Minimum of 3 UC ladder faculty, 2 (including Chair) must be in home department
• Additional members may be at departmental discretion

Departmental Committees Requirement

• Students must form a Ph.D. committee during their first year

Written Exams

Students are required to complete both written examinations within two years after enrolling in the Ph.D. program. Doctoral students, in consultation with their adviser and doctoral committee, chose two fields of study for their written exams. One will be a sub-discipline within ecology and/or evolution and the other may be another sub-discipline or may be in another discipline entirely but related to the development of the student's Ph.D. research. The student's reading and research in the period of foundation building will prepare them for the upcoming exams.

Written exams will test a student's understanding and their ability to work with material and ideas in a field at a level above that expected in an upper division undergraduate class. Preparation is done throughout the period of foundation building via graduate coursework, participation in seminars, and reading in advanced textbooks and independent reading. Students should plan to spend at least a few weeks of intense directed preparation immediately before the exam. Exams differ in structure and format (open vs. closed book, time limits, etc.), but they are all designed to test for the same, high level of understanding and performance.

If written exams are not completed within 2 years, the student will receive a reminder from the Graduate Advisor that they are falling off schedule. After 3 years, the student will receive a warning that they are off schedule and may be at risk of being dropped from the program if they do not complete the exams within the next year.

Students failing to successfully complete their written examinations within 4 years after enrolling in the Ph.D. program may be dropped from the program.

Students who fail a written exam must successfully complete the same or an alternative exam approved by their committee by the end of the following academic quarter. Students who fail two written exams will be dropped from the program. While failing an exam is unusual; conditional passes are fairly common. A conditional pass is given when the overall exam is satisfactory, but some aspect of it is weak. In such cases, the student will be expected do more reading, and then revise the weak section. Until this is done, the exam is considered incomplete.

To be completed within three years after enrolling in the Ph.D. program.

The oral exam is usually a broad based exam testing a student's knowledge and ability to work with material in the basic science areas relevant to their research. The oral exam committee will generally be the student's dissertation committee. In most cases, the oral exam will begin with a student's presentation of their dissertation proposal and the questioning may spin off that. However, a student should be prepared for the questioning to get very far afield from their specific proposal. Students should expect to find themselves questioned about things that they don't know the answer to, possibly including issues where no one knows the answer. Evaluating how a student handles uncertainty is an important part of evaluating whether they are ready to be advanced to candidacy for the Ph.D.

After passing the oral exam, a student will be "Advanced to Candidacy". That has two possible effects on a student's life. First, a student becomes eligible for doctoral candidate borrowing privileges at the Davidson Library. It also means that non-resident supplemental tuition will be waived for three years for international students.. Since EEMB students generally don't pay their own fees, this may not seem important, but whatever source is supporting them (departmental funds, grants, etc.) may benefit, by allowing resources to be used to support other students. Note, to advance to candidacy officially, you must file the Doctoral Form II with the Graduate Division and pay a fee (currently $50).

If the oral exam is not completed within 3 years, the student will receive a warning that they are falling off schedule, and that the exam must be completed within the next year or they may be placed on academic probation by Graduate Division. Students failing to successfully complete their oral examinations and advance to candidacy within 4 years will be placed on academic probation unless the Department petitions Graduate Division not to do so. This requires a solid justification.

Students who have not completed their oral exam and advanced to candidacy within 5 years after enrolling in the Ph.D. program may be dropped from the program.

Students who fail their oral examination are allowed to retake the exam once only. The second exam must be taken within one year.

During this period, the student is actively engaged in research including experiments, observations, analysis and writing. Students must manage their time wisely between demands for research, teaching, and public and professional service. It is important that students stay engaged with other graduate students for discussions and feedback, and with their adviser and the other members of their doctoral committee. The worst thing that a student can hear when they think they are finished with their research is "You need to have done…" The only way to ensure this doesn't happen is through regular interaction with the advisory committee.

Defense and Dissertation

The main thing a student needs to do after advancing to candidacy is to finish their dissertation. The official guidelines for format and filing requirements are available in the Guide to Filing of Theses and Dissertations. During the time a student is focusing on their research and writing, it is important that they stay in touch with their committee.

This is the length that the Department believes is a reasonable amount of time for a student to complete a Ph.D. In EEMB, normative time is 6 years following enrollment as an EEMB graduate student (in either a M.A. or Ph.D. track). If a student hasn't completed their Ph.D. within normative time, they lose the reduced fee benefit they may have received when they advanced to candidacy.

In accordance with university policy, students who have not completed their degree within 7 years of enrolling in the program must petition the Graduate Committee and the Graduate Dean to remain in the program. The petition must outline reasons for not completing the degree and an acceptable alternate completion schedule.

Students failing to complete their degree in 8 years from the time of their enrollment must retake their written examinations or prove in some equivalent way that they are fully up to date in their major field of interest. Simply completing the dissertation is not considered such proof, as a dissertation may be narrowly focused.

Students failing to complete the degree in 10 years from the time of their enrollment will be dropped from the program.

A final examination defending the dissertation will be made after the written dissertation is completed. Students may petition to have the final oral examination waived, and to give a public seminar of their dissertation research, in lieu. Petition forms to waive the final defense are provided to the Department by the Graduate Division and are available from the departmental office.

College of Biological Sciences

Department of Evolution and Ecology

The Department of Evolution and Ecology is a dynamic and collaborative community of faculty, students, post-doctoral researchers, and staff. Our mission centers around unraveling the intricate tapestry of evolution and ecology across diverse biological scales. From the evolution of genes and genomes to the diversification of species over time, we explore the fascinating interplay between life forms and their environments. Our research excellence spans these levels, delving into the structure, function, and biogeography of ecological communities. While our faculty members are committed to advancing scientific knowledge, they equally prioritize nurturing the next generation of scientists through quality teaching and mentorship. 

Undergraduate Programs

More information about CBS majors and minors

Graduate Groups

Facilities and programs.

Bodega Marine Lab

The Bodega Marine Laboratory, a leading research and education facility, serves society by providing science that informs solutions to complex environmental challenges in coastal ecosystems.

Center for Population Biology

The Center for Population Biology advances understanding of the fundamental ecological and evolutionary processes that control the origins and maintenance of biological diversity, at levels ranging from molecules to ecosystems.

Coastal and Marine Sciences Institute

The Coastal and Marine Sciences Institute, located on the UC Davis campus, conducts collaborative, cross-disciplinary research to address pressing challenges in coastal ocean sustainability.

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Marine Biology

Apply key principles from ecology, zoology, botany and genetics to examine marine life and investigate their biological relationship to their natural environments.

Guided by Australia’s leading marine researchers, you’ll be exposed to a variety of marine habitats and environmental conditions.

You'll gain extensive fieldwork experience at UQ’s Heron Island Research Station on the southern Great Barrier Reef and UQ’s Moreton Bay Research Station on North Stradbroke Island.

We'll show you how to apply a wide range of evidence-based ecological and environmental strategies to protect and sustainably manage vital marine organisms and ecosystems across the globe.

You'll be prepared for a broad range of career opportunities within research and education, aquaculture, wildlife conservation and ecotourism.

1 in Australia for environmental sciences

QS World University Rankings 2023

20 in the world for environmental sciences

Career possibilities

Our programs prepare you for your first job and beyond. Here are some of the careers you could be on your way to:

• Marine biologist
• Marine ecosystems researcher
• Marine environmental consultant
• Marine research officer
• Fisheries officer
• Fisheries research scientist

Graduate salary

Agriculture & environmental studies (undergraduate)

compared.edu.au

UQ runs one of the best Marine Biology programs in the country... Spending a week at Heron Island studying coral reef geology and collecting skin samples from dugongs in Moreton Bay are just two of the many highlights from studying at UQ.

My honours year allowed me to conduct research on major real-world issues that are currently affecting coral reef ecosystems on a global scale. This taught me how to think like a scientist, and create relevant research questions and studies on our most pressing issues.

Studying marine biology exposed me to coral reef research specific to the Great Barrier Reef and gave me the opportunity to spend a lot of time at the Heron Island Research station learning fundamental principles and processes of marine systems.

30 April - 1 May

Medicine Information Evening

1 May - 22 May

UQ's Sustainable Energy programs, Q&A webinars

Discover Postgraduate Public Health

3 pathways to engineering and computing at UQ

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Careers in computer science

Why study psychology?

How you'll learn.

Your learning experiences are designed to best suit the learning outcomes of the courses you choose.

• Work placements
• Overseas study
• Research experience
• Laboratory work

What you'll study

At UQ, subjects are called 'courses'. Here's a sample of the courses you could study:

• Fish, Fisheries and Aquaculture
• Living Oceans
• Global Challenges in Biology
• Marine Ecology and Conservation
• Ecology & Evolution of Marine Invertebrates
• Practical Marine Science

See courses and program structure

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Marine Biology

• College of the Environment
• University of Washington

New Career Resources Website from the College of the Environment

Looking for career resources for jobs in the environmental sciences?

The new Career Resources website  for College of the Environment students is now live on the  Dean’s Office webpage !

This new site is primarily geared for undergrads and contains information and resources to help you:

• find jobs, internships, and research opportunities
• prepare application materials
• explore graduate school and careers in academia

They also have a Career Services Events calendar   with upcoming workshops, programs, and other events to further your career goals. You can  subscribe to the calendar   for a weekly email update of upcoming events.

Bookmark their website for easy access as you look for opportunities during your time at the UW and beyond!

Congratulations to Charlie Thrift

Congratulations to Charlie Thrift for receiving a Graduate Research Fellowship Program (GRFP) award!

Charlie is studying insect conservation, community, and island ecology, co-advised by Hillary Young and Katja Seltmann.

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Hillary Young Lab • Ecology, Evolution, and Marine Biology • UC Santa Barbara © 2023 Regents of the University of California

Best Global Universities for Engineering in Russia

These are the top universities in Russia for engineering, based on their reputation and research in the field. Read the methodology »

To unlock more data and access tools to help you get into your dream school, sign up for the  U.S. News College Compass !

Here are the best global universities for engineering in Russia

Itmo university, tomsk state university, tomsk polytechnic university, lomonosov moscow state university, novosibirsk state university, saint petersburg state university, peter the great st. petersburg polytechnic university, moscow institute of physics & technology, national research nuclear university mephi (moscow engineering physics institute).

See the full rankings

• Clear Filters
• # 307 in Best Universities for Engineering  (tie)
• # 696 in Best Global Universities  (tie)
• # 364 in Best Universities for Engineering  (tie)
• # 587 in Best Global Universities  (tie)
• # 396 in Best Universities for Engineering  (tie)
• # 879 in Best Global Universities  (tie)
• # 632 in Best Universities for Engineering  (tie)
• # 355 in Best Global Universities
• # 809 in Best Universities for Engineering  (tie)
• # 579 in Best Global Universities  (tie)
• # 847 in Best Universities for Engineering  (tie)
• # 652 in Best Global Universities
• # 896 in Best Universities for Engineering  (tie)
• # 679 in Best Global Universities  (tie)
• # 902 in Best Universities for Engineering  (tie)
• # 475 in Best Global Universities  (tie)
• # 915 in Best Universities for Engineering  (tie)
• # 483 in Best Global Universities  (tie)

Victor Mukhin

• Scientific Program

Title : Active carbons as nanoporous materials for solving of environmental problems

However, up to now, the main carriers of catalytic additives have been mineral sorbents: silica gels, alumogels. This is obviously due to the fact that they consist of pure homogeneous components SiO2 and Al2O3, respectively. It is generally known that impurities, especially the ash elements, are catalytic poisons that reduce the effectiveness of the catalyst. Therefore, carbon sorbents with 5-15% by weight of ash elements in their composition are not used in the above mentioned technologies. However, in such an important field as a gas-mask technique, carbon sorbents (active carbons) are carriers of catalytic additives, providing effective protection of a person against any types of potent poisonous substances (PPS). In ESPE “JSC "Neorganika" there has been developed the technology of unique ashless spherical carbon carrier-catalysts by the method of liquid forming of furfural copolymers with subsequent gas-vapor activation, brand PAC. Active carbons PAC have 100% qualitative characteristics of the three main properties of carbon sorbents: strength - 100%, the proportion of sorbing pores in the pore space – 100%, purity - 100% (ash content is close to zero). A particularly outstanding feature of active PAC carbons is their uniquely high mechanical compressive strength of 740 ± 40 MPa, which is 3-7 times larger than that of  such materials as granite, quartzite, electric coal, and is comparable to the value for cast iron - 400-1000 MPa. This allows the PAC to operate under severe conditions in moving and fluidized beds.  Obviously, it is time to actively develop catalysts based on PAC sorbents for oil refining, petrochemicals, gas processing and various technologies of organic synthesis.

Victor M. Mukhin was born in 1946 in the town of Orsk, Russia. In 1970 he graduated the Technological Institute in Leningrad. Victor M. Mukhin was directed to work to the scientific-industrial organization "Neorganika" (Elektrostal, Moscow region) where he is working during 47 years, at present as the head of the laboratory of carbon sorbents.     Victor M. Mukhin defended a Ph. D. thesis and a doctoral thesis at the Mendeleev University of Chemical Technology of Russia (in 1979 and 1997 accordingly). Professor of Mendeleev University of Chemical Technology of Russia. Scientific interests: production, investigation and application of active carbons, technological and ecological carbon-adsorptive processes, environmental protection, production of ecologically clean food.   

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30 Best universities for Mechanical Engineering in Moscow, Russia

Updated: February 29, 2024

• Art & Design
• Computer Science
• Engineering
• Environmental Science
• Liberal Arts & Social Sciences
• Mathematics

Below is a list of best universities in Moscow ranked based on their research performance in Mechanical Engineering. A graph of 269K citations received by 45.8K academic papers made by 30 universities in Moscow was used to calculate publications' ratings, which then were adjusted for release dates and added to final scores.

We don't distinguish between undergraduate and graduate programs nor do we adjust for current majors offered. You can find information about granted degrees on a university page but always double-check with the university website.

1. Moscow State University

For Mechanical Engineering

2. Bauman Moscow State Technical University

3. National Research University Higher School of Economics

4. Moscow Aviation Institute

5. N.R.U. Moscow Power Engineering Institute

6. National Research Nuclear University MEPI

7. National University of Science and Technology "MISIS"

8. Moscow Institute of Physics and Technology

9. Moscow State Technological University "Stankin"

10. RUDN University

11. Moscow Polytech

12. Moscow State University of Railway Engineering

13. Finance Academy under the Government of the Russian Federation

14. Moscow Medical Academy

15. Russian State University of Oil and Gas

16. mendeleev university of chemical technology of russia.

17. Russian National Research Medical University

18. Plekhanov Russian University of Economics

19. National Research University of Electronic Technology

20. Moscow State Pedagogical University

21. Russian Presidential Academy of National Economy and Public Administration

22. State University of Management

23. Moscow State Institute of International Relations

24. Russian State Geological Prospecting University

25. russian state agricultural university.

26. New Economic School

27. Moscow State Technical University of Civil Aviation

28. Russian State University for the Humanities

29. Russian State Social University

30. Moscow State Linguistic University

Universities for Mechanical Engineering near Moscow

Engineering subfields in moscow.